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==Carnegie Stage 1 to 23== | [[File:Embryo stages 002 icon.jpg|link=Human Embryo Development Movie|100px]]

==Carnegie Stage 1 to 23== | <mediaplayer width='226' height='320'>file:Embryo stages 002.flv</mediaplayer>

...evelopment of Early Human Embryos: Observation of 90 Specimens at Carnegie stages 7 to 13= ...essed embryonic age and growth — was made for 90 human embryos at Carnegie stages 7–13 from healthy pregnancies.

...egie Collection]] normal and abnormal human embryo development during week 1. {{Carnegie stage 2 links}}

...at the external appearance of embryos in sequence from [[#Carnegie_Stages|1 to 23]]. It is not so important to memorise the dates, as they are only app Clicking the Carnegie stage numbers opens a page dedicated to describing that single stage and th

...to mesenchymal cell transition and folding of the embryonic disc. Carnegie stages 7 to 9 covers this period of human embryo development. ==Carnegie Stages==

{| border="1" cellpadding="10" | <Flowplayer height="300" width="226" autoplay="true">Embryo stages 002.flv‎</Flowplayer>

...many of his drawings and plates forming the main visual component of many Carnegie publications. There was a 1992 article on his artistic contribution to embr ...Hopkins in 1913, Didusch was appointed as its illustrator. He remained the Carnegie Illustrator until his death in 1955."

=Developmental Stages in Human Embryos= |+ '''Including a Revision of Streeter's "Horizons" and a Survey of the [[Carnegie Collection]]'''

...ult (M, G₁, S, G₂, M phase). With virtually no G₁ or G₂ phases this results in a reduction in cytoplasmic vo * Week 1, 2 - 3 days, size 0.1 - 0.2 mm

* Week 1, size 0.1 - 0.15 mm {{Carnegie stage 1 links}}

...external changes on the embryo. These stages are also shown in the movie "Carnegie stage human growth through embryonic period". Identify the key structures i | <html5media height="496" width="386">File:Embryo stages 003.mp4</html5media>

* Week 1, 4 - 5 days, size 0.1 - 0.2 mm {{Carnegie stage 3 links}}

...age or size. The human embryonic period proper is divided into 23 Carnegie stages. Criteria beyond morphological features include age in days, number of somi ...historical US Institute that historically categorised these developmental stages.

{{Carnegie stages}} ==Selected Embryo Stages==

[[Sensory - Hearing and Balance Development]] | [[Carnegie Embryos]] ...hilly 1987|link=Embryology History - Ronan O'Rahilly|Ronan O'Rahilly (1987 Carnegie Labs)]]

name the three stages of follicle development in the ovary 1. What factor do the synctiotrophoblast cells secrete to support the ongoin

--[[User:Z3218146|Julianna Lam]] 13:42, 17 September 2009 (EST)'''lab 1''' 1-what is the protein that the sperm binds to on the surface of the egg

'''[[Lab 1]]''' 1. What is the protein that the sperm binds to??? ZP3

'''Modern Notes:''' [[Carnegie stage 2]] | [[Week 2]] [[:Category:Carnegie Embryo 8190|'''Carnegie, No. 8190''']]

...embryology is by direct observation of histological sections at different stages of development. Even today the best way to observe anatomical development a ...d 22 images are available in at least two formats, an unlabeled set (image 1 - 49) and a labeled set (image 50 - 98) in a rostro-caudal sequence (head t

...k Hill]] 21:58, 5 August 2009 (EST) Where are your answers from Laboratory 1 Assessment? Attended lab--[[User:Z3187802|Vishnnu Shanmugam]] 13:05, 1 October 2009 (EST)

.... 292 p.</ref> There has been a recent photographic study of developmental stages in Chinook salmon (''Oncorhynchus tshawytscha'') embryonic development.<ref ...objectives of this study were to construct an index relating developmental stages of hatchery-reared fall Chinook salmon embryos to time and temperature (e.g

...l Stages In Human Embryos - Appendix 1|Appendix 1]] | [[1987 Developmental Stages In Human Embryos - Appendix 2|Appendix 2]] ==Appendix II - Index of Specimens of Stages 1-9==

| [[File:Week2_001 icon.jpg|left|300px|link=Carnegie stage 4#Stage 4 Animation]] * Week 1-2, size 0.1 - 0.2 mm

{{Carnegie Collection stage 22 table}} ...row, [[:Category:Carnegie Embryo 6832|No. 6832]]. Bottom row, [[:Category:Carnegie Embryo 8394|No. 8394]]. All views are at the same magnification.

== '''Lab 1 Questions''' == 1)What is the protein that sperm binds to on the surface?

==Carnegie Stages== Historic [[Carnegie Stages]] 10 to 13.

1) What is the protein that sperm binds to on the surface of the egg? 2) Name the 3 stages of follicle development in the ovary?

Lab 1 Questions 1. What is the protein that the sperm binds to on the surface?

'''Lab 1 Questions:''' ''Q2. Name the 3 stages of follicular development in the ovary:''

==Lab 1 Questions== ''1. What is the protein that the sperm binds to on the egg surface?''

Use the links to [[Carnegie stage 7]], [[Carnegie stage 8]] and [[Carnegie stage 9]] to see a number of different views of the human embryo in the thi [[Carnegie stage 7|Human embryonic stage 7]] occurs during week 3 between 15 to 17 day

...hn Rock|John Rock]] and included embryos that were incorporated into the [[Carnegie Collection]]. [[Carnegie Collection]] Embryo No. {{CE8004}} (stage 5b) from 1945 paper.<ref name=He

You have now been through the earliest stages of embryonic development. From the original simple trilaminar embryo all th ...s also divided into 23 stages (I do not expect students to identify embryo stages).

1- What is the protein that the sperm binds to on the surface? 2-Name the three stages of follicle development?

[[File:Week1 summary.jpg|thumb|Week 1 and 2 development]] {{Week 1 Links}}

=Lab 1 Questions= 1.What is the protein that the sperm binds to on the surface?

Lab 1 Q1: What is the protein that sperm binds to on the surface of the egg? ZP3 Lab 1 Q2: Name the three stages of follicle development in the ovary: Primordial, pre-antral and Graafian.

...sets were incorporated in 1972 into the [[Carnegie Collection]] (assigned Carnegie Nos. 10315-10434), but have since been returned to the University of Goetti | valign="bottom"|{{Embryo 1.6mm movie 1‎}}

...art rate are very similar between the 17th and 23rd Carnegie developmental stages make the baboon a promising surrogate of human pregnancy for investigations Genetic code: Translation table 1 (Standard)

| valign=top|All human and animal embryos go through very similar stages of early development. ...jor difference appears to be how long it takes to reach each of these same stages.

==Carnegie Institution of Washington Year Book 1952== ...om the year book.<ref>Carnegie Institution of Washington Year Book No. 51. Carnegie Institution Of Washington Washington, D. C. 1952</ref>

== 30/7/09 Lab 1 Questions == 1. What is the zona pellucida protein that binds spermatozoa to the oocyte su

See also [[Carnegie stage 12#Events|'''Carnegie stage 12 Events''']] {{Carnegie stage 12 links}}

...le:Week1 summary.jpg|thumb|alt=Week 1 and 2 development cartoon|300px|Week 1 and 2 development]] ...Events of Human Development during the first week ([[:Category:Week 1|week 1]]) following fertilization or clinical [[Gestational Age|gestational age]]

...p3|listen Part 5]] | [[:File:BGD2010-Embryo Lab 170510-605.mp3|download]] (1.8 Mb MP3 13:14) ...evelopment and look at other system organization compared with the earlier stages.

[[Carnegie stage 20]] '''Fig. 20-1.''' Photographs of three embryos belonging to stage 20. A slight bend is pr

...k Hill]] 21:59, 5 August 2009 (EST) Where are your answers from Laboratory 1 Assessment? 1. What factor do the synctiotrophoblast cells secrete to support the ongoing

See also [[Carnegie stage 14#Events|'''Carnegie stage 14 Events''']] * midbrain, nasal placode, lens pit, 1,2,3 pharyngeal arches, fourth ventricle of brain, 1st pharyngeal groove, he

== WEEK 2 LAB 1 == '''1)''' What is the protein that sperm bind to on the egg surface:

===Lab 1 Questions=== '''''1. What is the zona pellucida protein that binds spermatozoa to the oocyte su

...k Hill]] 21:58, 5 August 2009 (EST) Where are your answers from Laboratory 1 Assessment? ===Lab 1 Questions===

See also [[Carnegie stage 15#Events|'''Carnegie stage 15 Events''']] {{Carnegie stage 15 links}}

There are 2 Carnegie stages that show external embryo development during this week. File:Stage16_bf1.jpg|[[Carnegie stage 16|Stage 16]]

* Week 6 to 7 - [[Carnegie_stage_17|Stages 17 - 19]] - Retinal pigment is visible and the retinal fissure is largely * Week 7 - [[Carnegie_stage_18|Stages 18]] - Mesenchyme invades the region between the lens epithelium and the su

...-4377-2002-0&uniqId=330028653-2#4-u1.0-B978-1-4377-2002-0..00001-1 Chapter 1 - Introduction to the Developing Human] ...4377-2002-0..00002-3&isbn=978-1-4377-2002-0&uniqId=330028653-2#4-u1.0-B978-1-4377-2002-0..00002-3 Chapter 2 – First Week of Human Development]

'''Lab 1 Questions''' 1. What is the protein that the sperm binds to on the surface of the ova?

...ent species can also be classified by the same [[Carnegie Stage Comparison|Carnegie stage system]]. ...er than mouse embryos and easy to breed. Rat development is also generally 1 day behind from mouse.

[[1. An Early Ovum imbedded in the Decidua_(1908)|'''1. An Early Ovum Imbedded In The Decidua''']] ...e production a costly one, and we desire to express our obligations to the Carnegie Trust of the Scottish Universities for giving us a grant towards the expens

See also [[Carnegie stage 18#Events|'''Carnegie stage 18 Events''']] * Development indices: number of semicircular ducts (1-3) and length of the paramesonephric duct.

[[File:Carnegie_stages_species_comparison.jpg|thumb|300px|Carnegie stages species comparison]] ...ematic development. The table below also has detailed descriptions of each Carnegie stage as well as identifying embryo examples from different collections and

See also [[Carnegie stage 21#Events|'''Carnegie stage 21 Events''']] {{Carnegie stage 21 links}}

==Stage 1== [[Carnegie stage 1]]

...16:53, 12 November 2009 (EST) [[Movie - Stage 16 to 18 Face]] added fixed Carnegie stage links in week by week pages. ...]] ventricular septation model movies [[Movie - Ventricular Septum 01|Part 1]] | [[Movie - Ventricular Septum 02|Part 2]] | [[Movie - Ventricular Septum

...series [[Book_-_Contributions_to_Embryology|Contributions to Embryology]] (Carnegie Institution of Washington). :'''Links:''' [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]]

[[2009_Group_Project_1|Project 1 - Rabbit]] | [[2009_Group_Project_2|Project 2 - Fly]] | [[2009_Group_Projec ...detailed the short-tailed fruit bat (''Carollia perspicillata'') embryonic stages of development.

See also [[Carnegie stage 17#Events|'''Carnegie stage 17 Events''']] {{Carnegie stage 17 links}}

* "Stages" refer to the Carnegie stages of development. * Week 1 to Week 8 ({{GA}} 10)are considered the {{embryonic}} period of development

See also [[Carnegie stage 11#Events|'''Carnegie stage 11 Events''']] {{Carnegie stage 11 links}}

See also [[Carnegie stage 16#Events|'''Carnegie stage 16 Events''']] {{Carnegie stage 16 links}}

Endochondral ossification within the limb begins at Carnegie stage 18 and also occurs throughout embryo skeleton. This process is the re ==Ossification Stages==

{{Carnegie No.20 Header}} File:Streeter001.jpg|Figure 1

==Introduction To Stages 19-23== | For stages 19–23, Streeter devised a system of rating the development of selected st

===Embryo Stages and Events=== | [[File:Stage11_bf1c.jpg|200px|link=Carnegie stage 11]]

[[File:Stage11 sem100.jpg|thumb|Human Embryo (Carnegie Stage 11)]] [[File:Stage11_sem6.jpg|thumb|'''Axes''' - Human Embryo (Carnegie stage 11), 24 days, 13 somite pairs]]

===Embryo Stages and Events=== | [[File:Stage11_bf1c.jpg|200px|link=Carnegie stage 11]]

..., St. Louis University, Missouri until 1969 when he became Director of the Carnegie Laboratories of Embryology in Baltimore, Maryland. ...ogy History - Fabiola Müller|Fabiola Müller]] based upon studies of the [[Carnegie Collection]] of human embryos.

...uestion will be found on page 278. The drainage of the femoral plexus (fig. 1, rete femorale) is accomplished in two ways. A portion of the blood travers * 1 {{Ref-Senior1919}} pp. 67 and 88, also figs. 3 and 9C. (Unfortunately, the

''The norm should be established; embryos should be arranged in stages.''<br><br>[[Embryology History - Franklin Mall|Franklin P. Mall]] ==The Carnegie Collection==

...ryology-human-timeline Human Timeline] | [http://thebox.unsw.edu.au/video/1-minute-embryology-endoderm-development Endoderm] ...utu.be/VcpAL8TcQCk UNSW Foundations (2016) - Introduction to Embryology] a 1 minute introduction to finding the lecture. This video is also available fr

...]] [[File:Endoderm_002_icon.jpg|90px|link=Development_Animation_-_Endoderm 1]] ===Embryo Stages and Events===

| valign=top|[[File:Stage5 bf11L.jpg|alt=Human embryo Carnegie stage 5|400px]] | valign="top"| '''Facts:''' Week 1 - 2, size 0.1 - 0.2 mm

* [[#Embryonic Stages (Carnegie)|Embryonic Stages (Carnegie)]] * [[#Fetal Stages (Crown-Rump Length)|Fetal Stages (Crown-Rump Length)]]

[[File:Carnegie_Institute_of_Washington_logo.jpg|thumb|Carnegie Institute of Washington]] ...The papers documented not only early human development, using mainly the [[Carnegie Collection]] of embryos, but also that in animal models of development.

=The Chimpanzee Ovum In The Early Stages Of Implantation (About 10.5 Days)= Department of EmbryologyThe Carnegie Institution Of Washington, Baltimore, Maryland

...]] [[File:Endoderm_002_icon.jpg|90px|link=Development_Animation_-_Endoderm 1]] ===Embryo Stages and Events===

...age_22_image_217.jpg|thumb|300px|Human embryo brain meninges. ([[Week 8]], Carnegie stage {{CS22}})]] ...nal cord02.jpg|thumb|300px|Human embryo spinal cord meninges. ([[Week 8]], Carnegie stage {{CS22}})]]

...eiler Stages''' divides mouse development into 26 prenatal and 2 postnatal stages.<ref name=Theiler1972>{{Ref-Theiler1972}}</ref> '''Downs and Davies Stages''' is a more recent (1993) staging of gastrulating mouse embryos by morphol

[[File:Stage11 sem100.jpg|thumb|Human Embryo (Carnegie Stage 11)]] [[File:Stage11_sem6.jpg|thumb|'''Axes''' - Human Embryo (Carnegie stage 11), 24 days, 13 somite pairs]]

See also [[Carnegie stage 20#Events|'''Carnegie stage 20 Events''']] {{Carnegie stage 20 links}}

[[File:Human Carnegie stage 1-23.jpg|400px|Human Embryo, Carnegie stages 1-23]] ...n of these feature (relative to the adult position). Note that many of the stages have larger versions available and should be opened alone for best detailed

...le:Week1 summary.jpg|thumb|alt=Week 1 and 2 development cartoon|300px|Week 1 and 2 development]] ==Carnegie Stages==

...esenchymal cell transition and folding of the the embryonic disc. Carnegie stages 7 to 9 covers this period of human embryo development. :''From 1 layer of cells to 3 layers that define all tissues of the entire embryo.''

...heiler Stages''' divide mouse development into 26 prenatal and 2 postnatal stages and is based upon the publication '''The House Mouse: Atlas of Mouse Develo == Theiler Stage 1 ==

...gnancy embryos. This was the very first publication in the series from the Carnegie Institution of Washington called [[Book_-_Contributions_to_Embryology|Contr ...anklin Mall|'''Franklin P. Mall''']] Director of Department of Embryology, Carnegie Institution of Washington

* Users - 1.4 M * Sessions - 1.9 M

...du.au/Medicine/BGDlecture5.htm Lecture 9 Development of the embryo/fetus - 1] (12 May 10-11am, Clancy) ==Week 1==

'''Lab 1 Questions:''' '''1. What is the protein that the sperm binds to on the surface?'''

From The Department Of Embryology, Carnegie Institution of Washington, Baltimore, Maryland. ...a suitable suite of chambers for the contained membranous labyrinth in all stages of its development.

== Week 1 == [[File:Week1 summary.jpg|thumb|Week 1 and 2 development]]

# [[#1. Human Embryology History|'''Human embryology history''']] ==1. Human Embryology History==

{{Carnegie No.20 Header}} ...or studying the general question of growth of cartilage, for two reasons: (1) there are, on account of the intricacy of form of the labyrinth, many kind

....ncbi.nlm.nih.gov/pubmed/12807866?dopt=Abstract Development of the heart: (1) Formation of the Cardiac Chambers and Arterial Trunks] by Moorman et al. ( ...&Ausgabe=235779&ProduktNr=224197 Embryonic Cardiac Morphometry in Carnegie Stages 15-23, from the Complutense University of Madrid Institute of Embryology Hu

...o access during periods of development). Rat development is also generally 1 day behind that of mouse. (The table below gives details relating to the st ...y 20). A string of pearls was seen on days 8-11. The CR length ranged from 1.6  mm (day 11) to 26. 4 mm (day 20). A heart beat was first seen on day

...evelopment of these different portions of the coelom, as well as the early stages of the development of the diaphragm. ...th a transverse section at the level of the fourth cervical ganglion (fig. 1), the nerve was found arising from the anterior roots of the fourth cervica

# [[#1. Human Embryology History|'''Human embryology history''']] ==1. Human Embryology History==

...graphy also came into play, seen mainly associated with embryos within the Carnegie collection. ...s: the wax models are life-sized; the terracotta versions are reduced to a 1:3 scale. Together with the anatomical waxes in the Specola Museum in Floren

...s: black (C57BL/6J), agouti (CByB6F1/J) and albino (BALB/cByJ). When all 4 stages (proestrus, estrus, metestrus, and diestrus) need to be identified, vaginal ...MID19701181}} "In both sEHKO and GFP colonies, increases in body weight at 1 and 2 weeks after timed male exposure more reliably and consistently indica

...int Louis University, Saint Louis, Missouri, and Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland ...ic period of development (the first 7-8 weeks after fertilization) into 23 stages. Each stage was identified by a number of external and internal characteris

...o access during periods of development). Rat development is also generally 1 day behind that of mouse. (The table below gives details relating to the st ==Day 1==

{{Carnegie Collection stage 23 table}} ...er row, [[:Category:Carnegie Embryo 7425|No 7425]]. Lower row, [[:Category:Carnegie Embryo 4570|No. 4570]]. All views are at the same magnification.

...Heard (1891 – 1983) was one of the first to be hired, to the newly formed Carnegie Institute of Embryology in 1913, he was modeller who spent 42 years at the :'''Links:''' [[Carnegie Collection]] | [[Carnegie Stages]] | [[Embryology_History_-_Ziegler_Models|Ziegler Models]]

...cal characteristics of the miniature pig palate at different developmental stages were synchronously verified at the ultrastructural level. Our study provide * '''Axial differentiation and early gastrulation stages of the pig embryo'''{{#pmid:19683851|PMID19683851}} "Differentiation of the

Carnegie Stage 4 (stages 1-23 describing key steps in embryonic development) represents the beginning ...sentially covering the birth canal. This condition occurs in approximately 1 in 200 to 250 pregnancies.

...the [[Harvard Collection|Harvard Embryological Collection (H.E.C.)]] and [[Carnegie Collection]]. ...n of Anatomists, February, 1932; abstract, Anatomical Record, vol. 52, no. 1,pp. 2-3, February 25, 1932.

| [[File:Stage7-sem2.jpg|250px|link=Carnegie stage 7]] | [[File:Stage8_SEM1.jpg|250px|link=Carnegie stage 8]]

...pg|90px|left]] This 1949 historic paper by Hamilton describes the earliest stages of development following fertilisation. {{Week 1 Links}}

The points of special significance in regard to this material are: (1) the relatively longer persistence of the occlusion than has yet been descr ...regards an atresia of the esophagus in the human embryo as abnormal at all stages (p. 368).

[[Carnegie stage 4]] * Probably approximately 0.1-0.2 mm in diameter

{{Carnegie stage 10 links}} ==Carnegie Collection==

...p3|listen Part 2]] | [[:File:BGD2010-Embryo Lab 170510-602.mp3|download]] (1.9 Mb MP3 14:08) === Human Neuralation - Early Stages ===

[[Media:BGD2010-Embryo Lab 170510-601.mp3|listen Part 1]] | [[:File:BGD2010-Embryo Lab 170510-601.mp3|download]] (2.7 Mb MP3 20:39) ===Embryo Stages and Events===

==Carnegie Stage 4== ==Carnegie Stage 5==

[[Carnegie stage 3]] * Approximately 0.1 - 0.2 mm in diameter

...iscovered the "Yale" embryo and was a member of the Carnegie Institute's [[Carnegie Collection|embryology department]] research group from 1932 to 1971 when sh ...were carried out in the [[Carnegie Collection|Department of Embryology]], Carnegie Institution of Washington at Baltimore.) Subsequent checking of the {{monke

[[User:Z8600021|Mark Hill]] ([[User talk:Z8600021|talk]]) 16:11, 1 December 2016 (AEDT) Added educational presentation [[UNSW Medicine Learnin [[File:Kyoto940 stage21-07.jpg|thumb|150px|Kyoto embryo (940) [[Carnegie stage 21|stage 21]] histology]]

...[fertilization]] ({{GA}} week 3) and is described as [[Carnegie stage 2|'''Carnegie stage 2''']]. This stage is followed by formation of a cavity, the blastoco ...ion development and affected the expression levels of POU class 5 homeobox 1 (OCT-4) and SRY-related HMG-box gene 2 (SOX2), transcription factors necess

...n (8), Wagener (9) and Smyth (10), in {{rabbit}}s by Wagener(9) and Honore(1); in Dasyurus by O’Donoghue (12); in {{pig}}s by Corner (13), and Schmalt ...ve ova. It is possible that serial sections would have revealed more. Fig. 1 shows the follicle containing five ova, and is particularly noteworthy in t

{{Carnegie stage 1 links}} ...ittee for Research in Problems of Sex, National, Research Council, and the Carnegie Corporation of New York.

...nses The Cell Biology of the Senses] - [http://jcb.rupress.org/content/190/1/9.full The cell biology of hearing] July 12, 2010 . Based on data from the Carnegie embryo collection.<ref>{{Ref-O'Rahilly1963}}</ref>{{#pmid:6650859|PMID66508

==Carnegie Stages== [[File:Carnegie stages species comparison.jpg]]

...n]]. These human embryos are a mix of [[Carnegie stage 18|stage 18]] and [[Carnegie stage 19|stage 19]] embryos. ...Carnegie stage {{CS19}} {{CE144}}, 14 mm long; {{CE175}}, 13 mm long; and Carnegie stage {{CS18}} {{CE109}}, 10.5 mm long. In embryos of less length than no.

{{Carnegie stage 1 links}}

...''Links:''' [[Carnegie stage 10]] | [[:Category:Carnegie Stage 10|Category:Carnegie Stage 10]] | [[Week 4]] ...or approximately 47 days old. Beginning with stage xi, ovulation age 24 ± 1 days, Dr. Streeter carried the work forward through the remaining older gro

* [[Template:Carnegie_stages|Template:Carnegie_stages]]- menu link to each Carnegie stage page. ...ge table]] - table of Carnegie stages with brief descriptions and links to stages.

:'''Links:''' [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]] By '''George W. Corner''' with 2 plates and 1 figure, pp 61 - 66 (1922).

[[One_Minute_Embryology#Human_Development|'''1 Minute Embryology''']] | valign="bottom"|{{Human fertilization movie 1}}

==Carnegie Stage 12 to 14== * '''BA1-BA3''' - pharyngeal arches 1 - 3

...described the development of the vertebrae using human embryos from the [[Carnegie Collection]]. Epistropheus refers to cervical vertebra (C2) also named the [[Embryology History - Charles Bardeen|Charles Bardeen]] | [[Carnegie Collection]]

...hilly 1987|link=Embryology History - Ronan O'Rahilly|Ronan O'Rahilly (1987 Carnegie Labs)]] ...t study,<ref name=Weller1933>{{Ref-Weller1933}}</ref> used the following [[Carnegie Collection]] embryos: stage {{CS9}} (No. {{CE1878}}), {{CS10}} ({{CE391}};

[[One_Minute_Embryology#Human_Development|'''1 Minute Embryology''']] | valign="bottom"|{{Human fertilization movie 1}}

...tum in the 26 mm human embryo. Based on CRL this would make this about a [[Carnegie stage 22]] embryo in [[week 8]]. {{Carnegie stage 22 links}}

Carnegie Institution 013' Washington, Department Or Embryology ...llection of embryos, and of the sitting height of 48 fetal pigs in various stages of development; the procedure being done first upon the fresh specimens and

...cle, nerve, proliferation and hormone receptor markers. Key findings are: (1) The corpus cavernosum in males and females extends into the glans penis an ...tion: outer mean length= -5.514 + 0.622 *, and total mean length= -8.865 + 1.312 * (*= gestational weeks). The correlation coefficients were statistical

...es 3,368 - Page edits since Embryology was set up 46,327 - '''Views total 1,882,523''' ...es 3,284 - Page edits since Embryology was set up 45,016 - '''Views total 1,690,205'''

{{Carnegie No.20 Header}} this paper. In brief, they include: (1) the original condensation of the mesenchyme

{{Carnegie stage 6 links}} {{Carnegie stages}}

...e reported that with few exceptions the hands and feet fit into one of the stages ===Table 1===

...cluding factors II, VII, IX, and X, and the anticoagulant proteins C and S.1). It is contraindicated during pregnancy as crosses the placenta and increa ...cluding factors II, VII, IX, and X, and the anticoagulant proteins C and S.1). The exact teratogenic effect of fetal damage from warfarin therapy during

| valign="bottom"|{{Week 3 notochord 1 movie}} | valign="bottom"|{{Planar cell movement movie 1}}

...} ({{GA}} week 3) and is described as [[Carnegie stage 1|'''Carnegie stage 1''']]. This stage is followed by mitosis to form 2 blastomeres and then a so {{Carnegie stage 1 links}}

[[Special:Statistics|Statistics]]: Content pages - 1,351 Uploaded files - ...sion]] and [[Movie_-_Model_Embryo_to_128_Cell_Stage|Flash version]]. Added Carnegie collection [[Carnegie_stage_8#Carnegie_Collection|stage 8 images]].

[[Week 1]] | [[Week 2]] | [[Zygote]] | [[Morula]] | [[Blastocyst]] | [[Implantation] [[File:Week1_summary.jpg|thumb|450px|Week 1 and 2 Development (see [[Week 1 Movie]])]]

Although many human embryos of a presumed age of 15-17 days showing early stages of the primitive streak have been reported in the literat11re, most of them ...d was more normal than previously, lasting only 5 days; coitus occurred on 1, 5 and 13 (or 14) June. From the histological features of the embryo, it ap

...umb|300px|alt=Human embryonic cornea|Human embryonic cornea ([[Week 8]], [[Carnegie stage 22]])]] ...px|alt=Human embryonic cornea|Human embryonic cornea detail ([[Week 8]], [[Carnegie stage 22]])]]

...est correlation exists between cyemic and chorionic size, during the early stages of development, when the conceptus probably is still more nearly spherical. {| border=1

...nt is much less clear than in the limb. Here, by removing Shh at different stages during GT development in mice, we demonstrate a continuous requirement for Three main stages during development, mesonephric/paramesonephric duct changes are one of the

...riptions. These stages are also shown in the movie [[Embryonic_Development|Carnegie stage human growth through embryonic period]]. # Identify the early stages of embryonic development

| Primitive node cilia (human [[Carnegie stage 8|stage 8]]) ...ed that zinc finger transcription factors zinc finger transcription factor 1 (znfl1s) are expressed in the tailbud and axial mesoderm in zebrafish. Howe

[[Carnegie stage 2]] * Approximately 0.1 - 0.2 mm in diameter

...in the available series of models illustrating the human heart at various stages in its development. As these gaps become narrower, we are confronted with t ...basis of interpolations between more carefully observed earlier and later stages, and (3) the heart was particularly well preserved to demonstrate internal

...portion of the pancreas. The study include embryos and fetuses from the [[Carnegie Collection]]. The work falls into two categories: (1) the study of the embryonic normal {{pancreas}}, and (2) the study of the p

...the earlier months are even more rare. Streeter (’19) reported that in the Carnegie collection there were only forty—three specimens, of which all but two we

...' - Wednesday 11 am -1 pm in Anatomy Labs, Biosciences Building D26, Level 1 (Weeks 4 and 7 in Wallace Wurth 120). ...dissect and investigate life chicken embryos, determine the developmental stages, and annotate the structures.

...' - Wednesday 11 am -1 pm in Anatomy Labs, Biosciences Building D26, Level 1 (Weeks 4 and 7 in Wallace Wurth 120). ...dissect and investigate life chicken embryos, determine the developmental stages, and annotate the structures.

...4}}. The appearance of the upper and lower limb buds are more like a later Carnegie stage {{CS16}} embryo occurring in week 6, 37 - 42 days, CRL 8 - 11 mm. ...pe, comparative size and arrangement is indicated in the accompanying Fig. 1.

| valign="bottom"|{{Week 3 notochord 1 movie}} | valign="bottom"|{{Mesoderm migration movie 1}}

...king under the auspices of the Department of Experimental Evolution of the Carnegie Institution of Washington, at Cold Spring Harbor, Long Island. To the Direc ...thesis constructed by reasoning backward from the observed anatomy of late stages, chiefly from the dispo sition of the foetal membranes at birth in cases of

[[File:Human Stage14 neural02.jpg|thumb|Human Embryo CNS ([[Carnegie stage 14|stage 14]]) showing cranial nerve development]] ...Brn3b(KO) RGC axons show correct but delayed pathfinding during the early stages of embryonic development. However, loss of Brn3b does not affect the anatom

=The History of the Earliest Stages in the Human Clavicle= ...girdle complexes in the frog and man, for which comparison examine figures 1 to 3.

...of the human embryo between Carnegie stage 19 to 23 in week 8 using the [[Carnegie Collection]] embryos. {{Carnegie stage 19 links}}

...e:Mark_Hill.jpg|50px|left]] This historic 1945 paper by Hertig describes [[Carnegie Collection]] early human extra-embryonic development in [[week 2]]. The his ...d of Obstetrics, Harvard Medical School, and the Department of Embryology, Carnegie Institution of Washington, Baltimore.

...s from week 3 to week 8 ({{GA}} week 5-10) and is divided into 23 Carnegie stages of embryonic development. There will also be a brief introduction to fetal ...man_Embryo|'''1 Minute Embryology''']] | [https://thebox.unsw.edu.au/video/1-minute-embryology-human-embryo UNSW theBox]

University of Chicago Embryo H279 was added to the Carnegie Collection as Embryo {{CE3709}}. [[Carnegie Collection]] Embryos used in this study {{CE836}}, {{CE3709}}, {{CE1878}} a

[[File:HillFlorian1931a fig01.jpg|thumb|400px|'''Fig. 1.''' Graphic reconstruction of embryonal area.<br>Cl. Cloacal membrane; Pr.s ...om the more caudal portion of the head-process in two important respects: (1) it is accompanied on both sides by thickened bands of mesoderm which are i

...c) pregnancy embryos. This was an early publication in the series from the Carnegie Institution of Washington called [[Book_-_Contributions_to_Embryology|Contr :" In the early stages of development we can determine by the extent of the exocoelom and its cont

Department Of Embryology, Carnegie Institution Of Washington, And The Departments Of Obstetrics And Radiology, ...the intervillous space by endometrial branches of the uterine veins (Fig. 1). Exchange of metabolites between mother and fetus takes place across the t

[[Image:Stage9sm.jpg|thumb|Carnegie stage 9 showing somite formation]] [[Image:Stage 9 SEM1.jpg|thumb|Carnegie stage 9 scanning electron microscope image showing somite formation]]

* embryo may develop through early stages, can erode through the uterine horn and reattach within the peritoneal cavi Carnegie Stage 4 (stages 1-23 describing key steps in embryonic development) represents the beginning

...day 50). Its development and position was historically used to stage late Carnegie embryos. By '''Ellen B. Finley''' with 2 plates, 1 text-figure, pp 155-161 (1923).

...day 50). Its development and position was historically used to stage late Carnegie embryos. By '''Ellen B. Finley''' with 2 plates, 1 text-figure, pp 155-161 (1923).

[[Carnegie stage 9]] * Approximately 1.5–2.5 mm

=== Group 1 === ...p_Project_1|Group Project 1 - Rabbit]] | [[Talk:2009_Group_Project_1|Group 1 Discussion]] | [[Rabbit Development|UNSW Embryology - Rabbit]]

...l Stages In Human Embryos - Appendix 1|Appendix 1]] | [[1987 Developmental Stages In Human Embryos - Appendix 2|Appendix 2]] ==Appendix 1 - Embryos In The Carnegie Collection==

===Carnegie Stages - Eye=== The following data is from a study of human embryonic carnegie stages{{#pmid:7364662|PMID7364662}} and other sources.

=Week 1 to 3= ...ult (M, G₁, S, G₂, M phase). With virtually no G₁ or G₂ phases this results in a reduction in cytoplasmic vo

...velopment, in Humans from week 3 to week 8 and is divided into 23 Carnegie stages of embryonic development. There will also be a brief introduction to fetal * Carnegie stages illustrate external development

...- Ernest Frazer|J. Ernest Frazer]]. Human embryo of 23 somites occurs at [[Carnegie stage 12]] in [[Week 4]]. {{Carnegie stage 12 links}}

...age11 sem21.jpg|thumb|300px|Human embryo neural crest cells ([[Week 4]], [[Carnegie stage 11|stage 11]])]] ...dditional transcription factors Pax7 and AP-2alpha. Importantly, while HNK-1 labels few migrating NCCs, p75(NTR) labels a large proportion of this popul

[[Special:Statistics|Statistics]]: Content pages - 1,745 Uploaded files - 8,388 Page edits since Embryology was set up - 113,178 ...y Models]] page with information about [[Embryology_Models#Carnegie_Models|Carnegie Models]].

Early stages of mammalian development may readily be obtained from the albino rat (''Mus ...es, spaced at intervals of about 18 hours, resulting in 2-, 4-, and 8-cell stages occur during transit through the oviduct. During the foiu-th segmentation t

...psule. These human embryos are [[Carnegie Embryos]] and fetuses from the [[Carnegie Collection]]. Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland

...age11 sem21.jpg|thumb|300px|Human embryo neural crest cells ([[Week 4]], [[Carnegie stage 11|stage 11]])]] ...dditional transcription factors Pax7 and AP-2alpha. Importantly, while HNK-1 labels few migrating NCCs, p75(NTR) labels a large proportion of this popul

| valign=top|All human and animal embryos go through very similar stages of early development. See also [[K12 Human and Other Animal Development|Hum {| border=1

[[Carnegie stage 15]] (figs. 15-1 to 15-3, and 16-4)

...6 surface hillocks (auricular hillocks), three on each of pharyngeal arch 1 and 2. ...s, from MRI data was traced to evaluate the movement between the different stages with identical magnification. Two different sets of reference axes were sel

...ng the ""Biggart" [[Carnegie stage 7]] and the younger "Macafee" embryo [[Carnegie stage 5]]. {{Carnegie stage 5 links}}

...mated age of 25 days would make this a [[Carnegie stage 10|stage 10]] or [[Carnegie stage 11|stage 11]] embryo. 100 diameters, Spencer, 1/2 in. Zeiss projection ocular.

| valign="bottom"|{{Neural Tube Closure 1 movie}} | valign="bottom"|{{Neural Crest movie 1}}

...e time elements thus appear to have been established by previous workers, (1) that the loss of nuclei by the red blood cells begins to be evident during ...rom the beginning of the 5th Week (5 mm) to the 24th week (220 mm) (tables 1 and 2) and included all of the material in the collection which was suffici

[[File:Stage14 stomach.jpg|thumb|Developing stomach mid embryonic period (Carnegie stage {{CS13}})]] ...w link to larger cross-sections of the mid-embryonic period (end week 4) [[Carnegie stage 13|stage 13]] embryo starting just above the level of the stomach and

{{Mitosis movie 1}} {{Mitosis movie 1 links}}

Carnegie Institution of Washington - Year Book No. 38 July 1, 1938 - June So, 1939

...s University|Johns Hopkins University Collection]] that later became the [[Carnegie Collection]]. Carnegie Embryos {{CE2}}, {{CE109}}, {{CE144}}, {{CE145}}

.../lectopia.telt.unsw.edu.au/lectopia/lectopia.lasso?ut=90&id=138299 Lecture 1 - Course Introduction] ...-4377-2002-0&uniqId=330028653-2#4-u1.0-B978-1-4377-2002-0..00001-1 Chapter 1 - Introduction to the Developing Human]

...reeter is a description of early {{twinning}}. The embryo is classified as Carnegie Embryo No. {{CE1522}}. Department of Embryology, Carnegie Institution of Washington

...-4377-2002-0&uniqId=330028653-2#4-u1.0-B978-1-4377-2002-0..00001-1 Chapter 1 - Introduction to the Developing Human] ...4377-2002-0..00002-3&isbn=978-1-4377-2002-0&uniqId=330028653-2#4-u1.0-B978-1-4377-2002-0..00002-3 Chapter 2 – First Week of Human Development]

File:Spiegel1626_table05fig1.jpg|Table 5 Fig. 1 Placenta Fetal Side | bgcolor="white"|Obstetric models life-size wax models and scaled (1:3) terracotta models, examples of techniques for teaching medicine and obst

[[Carnegie Collection]]: {{CE45}}, {{CE75}}, {{CE86}}, {{CE95}} ...e work complete it will be necessary to study more embryos at intermediate stages between the five given here, and others from the third montli up to the adu

...escribed embryo measured 11 mm CRL, probably corresponding to [[week 6]] [[Carnegie stage 16]]. A reconstructed model was also illustrated in the paper. '''Modern Pages:''' [[Carnegie stage 16]] | [[Week 6]]

* First Trimester (1 - 12 weeks) - embryonic and early fetal ...scientifically to describe the human conceptus at both embryonic and fetal stages of development.

...Surgeons’ Hall, Edinburgh ; Hon. Fellow, Ameriean Gynceeological Society; Carnegie Research Fellow I shall therefore consider: (1) The development of the round ligament and of the mammary pouch; and (2) Th

...ine editor has replaced the traditional Roman numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

Carnegie Institution of Washington, Department of Embryology ...detail in the near future in Contributions to Embryology, published by the Carnegie Institution of Washington.

...s of how the definitive GIT tube is formed and then at the overview of the Carnegie stage {{CS22}} embryo GIT from one end to the other. The tract now has a di ...d also be comparing the GIT appearance with the earlier embryonic {{CS13}} Carnegie stage.

[[File:Stage11 sem100.jpg|thumb|300px|Human embryo ([[week 4]], [[Carnegie stage 11]]) Somites]] ...}. In humans, the first somite pair appears at day 20 and adds caudally at 1 somite pair/90 minutes until on average 44 pairs eventually form.

...are differentially targeted in the CVG compared to the other tissues from stages 13-15, and we performed gene set enrichment analyses to determine different ...mouse Primordial Germ Cells (PGCs) and gonadal somatic cells at embryonic stages: E11.5, E12.5 and E13.5. Differential expression analyses revealed differen

| {{Primordial germ cell migration 1}} * This looped animation shows the 3 stages of kidney development (pink) in relation to the hindgut region between Week

#REDIRECT [[Lecture - Week 1 and 2 Development]] =Week 1 and 2=

[[File:Human Carnegie stage 10-23.jpg|thumb|300px|Carnegie Embryos]] ...collection numbering also incorporated the Blechschmidt embryo collection (Carnegie Nos. 10315-10434 ) in 1972, the collection embryos have now been returned t

...to the whole embryo, so that it seemed preferable to determine equivalent stages in the mouse and human on the basis of the development of as many embryolog ...minated from detailed study. Embryos admitted to study are listed in table 1.

[[Lecture_-_2014_Course_Introduction|Lecture 1]] - [https://lectures.unsw.edu.au/ess/echo/presentation/e7ad0e2e-7a04-4719- {{Human development movie 1}}

...nd condition of the tubules in the mesonephroi of human embryos at various stages from about 30 to 40 mm. ...xcept that the older embryos have proportionately more tubules in the last stages of the dedifferentiation.

...with EFIC and magnetic resonance images from a study of 52 human fetuses (Carnegie stage 13-23). Time course of atrial, ventricular, and outflow septation wer | valign="bottom"|{{Heart ventricular septum model 1}}

[[Carnegie stage 8]] * Approximately 1–1.5 mm in greatest length

...t Of Anatomy, University Of Virginia Communicated By A. G. Mayer, December 1, 1916 ...k. Aortic cell clusters would seem to be a common feature of certain early stages of vertebrate development.

...r ear. These human embryos are [[Carnegie Embryos]] and fetuses from the [[Carnegie Collection]]. Department of Embryology, Carnegie Institution of Washington, Johns Hopkins Medical School, Baltimore, Marylan

...tage8_bf9.jpg|thumb|300px|Human notochordal process and notochordal canal (Carnegie stage {{CS8}})]] ...ly roles in development and later roles in patterning surrounding tissues. 1. Mechanical, influencing the folding of the early [[E#embryo|embryo]]; 2. M

[[File:Stage_22_image_085.jpg|thumb|300px|Developing Human Spleen ([[Carnegie stage 22|stage 22]])]] ...uman embryonic period'''{{#pmid:25403423|PMID25403423}} "Between Carnegie stages (CSs) {{CS14}} and {{CS17}}, the spleen was usually recognized as a bulge i

...magnetic resonance imaging. The cartilaginous femur was first observed at Carnegie stage 18. Major anatomical landmarks were formed prior to the initiation of ...ole-genome microarrays to identify genes with differential expression at 5 stages of limb development (E9.5 to 13.5), during fore- and hind-limb patterning."

[[File:Adult mesentery 1.jpg|thumb|alt=Sagittal view of female abdominal cavity|Sagittal view of abd ...and 13'''{{#pmid:26995337|PMID26995337}} "We selected 37 human embryos at Carnegie Stage (CS) {{CS11}}-{{CS13}} (28-33 days after fertilization) and three-dim

...storic 1944 paper by Davies describes early human development in week 2 ([[Carnegie stage 5]]). {{Carnegie stage 5 links}}

...was lower (P ≤ 0.05) at day 30 than at day 23 (464.8 ± 16.1 vs.596.4 ± 28.1, respective" {{placenta}} ...and times to delivery PO being 4.3 +/- 1.6 and 64.3 +/- 0.3 days, 4.0 +/- 1.4 and 66.3 +/- 0.4 days, and 2.5 and 68 days for IUAI at 6, 7 and 8 days, r

...alt=Developing Human Thymus (Carnegie stage 22)|Developing Human Thymus ([[Carnegie stage 22|stage 22]])]] ...d the protease A disintegrin and metallopeptidase with thrombospondin type 1 motif 3 (ADAMTS3)."

...d are derived from pharyngeal arches 1 - 6 with the face forming from arch 1 and 2 and the frontonasal prominence. Each arch contains similar Arch comp ...nd_Palate|'''1 Minute Embryology''']]<br>[https://thebox.unsw.edu.au/video/1-minute-embryology-face-and-palate UNSW theBox]

...ingle chorionic sac which contained twin embryos, each 12.3 mm. long (fig. 1). There is a common yolk sac from which distinct yolk stalks arise near tog ...to describe it. He has also furnished the accompanying photographs (figs. 1 to 3).

...ingle chorionic sac which contained twin embryos, each 12.3 mm. long (fig. 1). There is a common yolk sac from which distinct yolk stalks arise near tog ...to describe it. He has also furnished the accompanying photographs (figs. 1 to 3).

...pg|90px|left]] This 1943 historic paper by Hamilton describes the earliest stages of development following fertilisation. * Embryo 1 - 17th day of the menstrual cycle, ovulation probably occurred less than 12

...d are derived from pharyngeal arches 1 - 6 with the face forming from arch 1 and 2 and the frontonasal prominence. Each arch contains similar Arch comp * Know the stages and structures involved in the development of the face.

[[Carnegie stage 10]] * Approximately 1.5–3 mm in length

[[Media:2015ANAT2341 Lecture 1 - Course Introduction.pdf|Lecture - Print PDF]] | valign="bottom"|{{Human fertilization movie 1}}

Carnegie Embryological Laboratory, Baltimore, Maryland ...the period of its inauguration, scant attention was given to the embryonic stages and as little to the ear of other animals. It is therefore not surprising t

| valign="bottom"|{{Neural Tube Closure 1 movie}} | valign="bottom"|{{Neural Crest movie 1}}

{{Carnegie stage 13 links}} {{Carnegie stage 14 links}}

...nin gene-related peptide (CGRP, nociceptive responses), vanilloid-receptor-1 (VR1, polymodal nociceptors) and calretinin (calcium signalling) cell immun '''Related Movies:''' [[Chicken Neural Crest Migration Movie 1|Migration 01]] | [[Chicken Neural Crest Migration Movie 2|Migration 02]] |

==1. Introduction== ...results primarily from the lack of adequate series of successive embryonic stages hitherto available for study. In the present paper, which is based upon the

...evelopment of the {{thyroid}} using several early embryos from the later [[Carnegie Collection]]. ...organ divided is a vesicle or is a solid body, although none of them found stages in which the thyroid was dividing. Remak placed the origin of the thyroid a

...ine editor has replaced the traditional Roman numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

Department of Embryology, Carnegie Institution of Washington ...available for this purpose has made it possible to fill out the essential stages in the development of these veins with considerable completeness. The work