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:'''Links:''' [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington Series]] | [[Endocrine - Parathyroid Development| ===Table 1===

{{Human Fertilization Movie 2 frame table}} ...a]] [[Category:Zygote]] [[Category:Carnegie Stage 1]] [[Category:Carnegie Stage 2]]

{{Carnegie stage 3 links}} | [[File:Human embryo day 5.jpg|Stage 3 Day 5]]

|+ '''Carnegie Stage 23 - Central Nervous System''' ...nside the Kyoto embryo ([[Carnegie stage 23|stage 23]], [[week 8]]). The [[Stage 23 MRI Movie 1|labelled sagittal head section movie]] show further detail.

{{Human Fertilization Movie 1 frame table}} ...a]] [[Category:Zygote]] [[Category:Carnegie Stage 1]] [[Category:Carnegie Stage 2]]

...ithelial to mesenchymal cell transition and folding of the embryonic disc. Carnegie stages 7 to 9 covers this period of human embryo development. ==Carnegie Stages==

{{Carnegie stage 2 links}} {{Carnegie stages}}

|+ '''Carnegie Stage 23 - Sagittal''' ...ittal MRI scan (left to right) through a Kyoto embryo ([[Carnegie stage 23|stage 23]], [[week 8]]). The movie starts on the lefthand side of the embryo and

...he Carnegie stage numbers opens a page dedicated to describing that single stage and the associated developmental events. ...detailed descriptions which can be viewed in a week by week format, by the Carnegie stages or integrated into a [[Timeline_human_development|'''Timeline of hum

[[Carnegie stage table]]

{{Carnegie stages}} ! colwidth=100px|Carnegie<br>Stage

...n either age or size. The human embryonic period proper is divided into 23 Carnegie stages. Criteria beyond morphological features include age in days, number ...an take from as little as 10 days in chickens to nearly 60 days in humans. Carnegie is the name of a historical US Institute that historically categorised thes

[[Carnegie stage 13 - serial sections|Carnegie Stage 13 Embryo]] the outer dark line in the sections is the '''amniotic membrane ===Table Legend===

:'''Links:''' [[Carnegie Stage 8 - "Dobbin" Embryo|"Dobbin" Embryo (stage 8)]] | [[Embryology History - James Hill|James Hill]] | [[Echidna Developme ==Stage 8 Embryo==

[[Sensory - Hearing and Balance Development]] | [[Carnegie Embryos]] ...hilly 1987|link=Embryology History - Ronan O'Rahilly|Ronan O'Rahilly (1987 Carnegie Labs)]]

See also [[Carnegie stage 12#Events|'''Carnegie stage 12 Events''']] * Caudal neuropore closes during this stage.

* "Stages" refer to the Carnegie stages of development. | <center>'''Stage'''</center>

...ombination in the zygote, generally fail to develop through the blastocyst stage to hatch and implant in the second week of development. These can occur wit :[[Carnegie_stage_table|'''Carnegie Stages''']]: [[Carnegie_stage_1|1]] | [[Carnegie_stage_2|2]] | [[Carnegie_s

Use the links to [[Carnegie stage 7]], [[Carnegie stage 8]] and [[Carnegie stage 9]] to see a number of different views of the human embryo in the third wee ==Stage 7==

...- good for what studies in rabbit embryo has been used for. And has a good table for embryological stages!! 1. What period of human development (in weeks) do the 23 Carnegie stages cover? 8 weeks

See also [[Carnegie stage 13#Events|'''Carnegie stage 13 Events''']] {{Carnegie stage 13 links}}

| valign="bottom"|{{Embryo stage movie 1}} Human blastocyst week 1 movies, 3 above movies together in single table.

...ecting surface and internal development. [[:File:Stage13_sem1.jpg|Carnegie stage 13]] | valign="bottom"|{{Stage 23 MRI movie 5}}

==Carnegie Stages== Historic [[Carnegie Stages]] 10 to 13.

...and table of [[Fetal_Development#Carnegie_Fetal|Carnegie embryos at fetal stage]]. ...dded video recordings of 2016 lectures on [[Template:Science Lecture movie table|Neural Crest, Head and Musculoskeletal development]].

* Stage 13 - Week 4, 26 - 30 days, 3 - 5 mm, Somite Number 21 - 29 The table below shows a thumbnail view of each section and a brief description of rel

==Stage 21== {{Carnegie Collection stage 21 table}}

See also [[Carnegie stage 14#Events|'''Carnegie stage 14 Events''']] {{Carnegie stage 14 links}}

==Stage 22== {{Carnegie Collection stage 22 table}}

There are 2 Carnegie stages that show external embryo development during this week. File:Stage16_bf1.jpg|[[Carnegie stage 16|Stage 16]]

| [[File:Stage7-sem2.jpg|250px|link=Carnegie stage 7]] | [[File:Stage8_SEM1.jpg|250px|link=Carnegie stage 8]]

==Carnegie Stages== Historic [[Carnegie Stages]] 1 to 4.

Human embryonic stage 9 occurs during week 3 between 19 to 21 days. ...tes have begun to form and number between 1 to 3 somite pairs during this stage.

==Human Carnegie Stages== | Carnegie stages are named after the famous USA institute which began collecting and

==Carnegie Stage 12 to 14== [[Carnegie_stage_11|Stage 11]]

...es can also be classified by the same [[Carnegie Stage Comparison|Carnegie stage system]]. {{Mouse Rat Pig table}}

...ate:Carnegie_stages|Template:Carnegie_stages]]- menu link to each Carnegie stage page. * [[Template:Carnegie stage table|Template:Carnegie stage table]] - table of Carnegie stages with brief descriptions and links to stages.

==Stage 1== [[Carnegie stage 1]]

See also [[Carnegie stage 16#Events|'''Carnegie stage 16 Events''']] {{Carnegie stage 16 links}}

...pment). Rat development is also generally 1 day behind that of mouse. (The table below gives details relating to the staging of rat development). * Standard Witschi Stage 1, day 1 , Size (mm) 0.07

| [[File:Human embryo day 2.jpg|200px|Stage 2 Day 2]] | [[File:Human embryo day 3.jpg|200px|Stage 2 Day 3]]

See also [[Carnegie stage 20#Events|'''Carnegie stage 20 Events''']] {{Carnegie stage 20 links}}

This early week 4 embryonic stage shows key features of heart tube and neural plate to tube development. See also [[Carnegie stage 11#Events|'''Carnegie stage 11 Events''']]

See also [[Carnegie stage 21#Events|'''Carnegie stage 21 Events''']] {{Carnegie stage 21 links}}

See also [[Carnegie stage 18#Events|'''Carnegie stage 18 Events''']] * Development indices: number of semicircular ducts (1-3) and length of the paramesonephric duct.

[[File:Carnegie_stages_species_comparison.jpg|thumb|300px|Carnegie stages species comparison]] ...velopment. The table below also has detailed descriptions of each Carnegie stage as well as identifying embryo examples from different collections and the p

Endochondral ossification within the limb begins at Carnegie stage 18 and also occurs throughout embryo skeleton. This process is the replacem * '''Stage 1''' - non-ossified epiphysis

See also [[Carnegie stage 17#Events|'''Carnegie stage 17 Events''']] {{Carnegie stage 17 links}}

Department of Embryology, Carnegie Institution of Washington, Baltimore ...study of this sort. Pertinent information for each embryo is presented in table 1, in which crown-rump length has been corrected in accordance with Streete

See also [[Carnegie stage 15#Events|'''Carnegie stage 15 Events''']] ...upper limb bud, mesonephric ridge, lower limb bud, umbilical cord Labelled Stage 15

! style="background:lightsteelblue" width="100" | '''Theiler Stage''' | Theiler Stage 1

==Carnegie Stage 12 to 14== [[Carnegie_stage_11|Stage 11]]

...int Louis University, Saint Louis, Missouri, and Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland ...us amphibians and notably the {{chick}} (Hamburger & Hamilton, 1951). Each stage is characterized by a number of external, morphological criteria, and it is

{{Carnegie stage 10 links}} ...membrane has been removed. Top embryo is an early stage 10, bottom is late stage 10.

This page is an out of date historic page (2004) table of contents by page format of the original [http://embryology.med.unsw.edu. ...find related content. These pages will remain ''frozen'' at their mid-2009 stage and still available as a DVD version. All updates will be added to '''2009'

| colspan=2|{{Human Fertilization Movie 1 frame table}} | [[File:Human embryo day 2.jpg|200px|Stage 2 Day 2]]

...various internal structures in order to assign the specimen precisely to a stage. ...not only the stage (Table 19-1) but even the relative position within that stage.

...2014 (EST) Added [[Model Embryo 7.5mm Movie 1|Human Embryo 7.5mm model]] (Stage 15) from the [[Blechschmidt Collection]]. Added draft historic textbook [[B ...tage_13#Hill_Collection|Stage 13]] and [[Carnegie_stage_16#Hill_Collection|Stage 16]] from [[Hill Collection]].

==Carnegie Stages== Historic [[Carnegie Stages]] 5 and 6.

==Stage 22 Spinal Cord== ...?zoom=4&lat=-7235.19233&lon=4189.80767&layers=B vertebral body] (cartilage stage)

| valign=top|[[File:Stage5 bf11L.jpg|alt=Human embryo Carnegie stage 5|400px]] * The '''conceptus''' completes implantation during this stage.

| valign="bottom"|{{Embryo stage movie 1}} His's Normentafel (Normal Table)

...1993 Downs and Davies staging of the mouse embryo included in each Theiler stage (Staging of gastrulating mouse embryos by morphological landmarks in the di == Theiler Stage 1 ==

{{Dobbin table}} {{Carnegie stage 8 links}}

| valign="bottom"|{{Embryo stage movie 1}} His's Normentafel (Normal Table)

==The Carnegie Collection== ...ollection of human embryos, begun in 1887, later became the basis of the [[Carnegie Collection]] (Mall and Meyer, 1921). Mall (1913) stated his indebtedness to

...an embryology page for [[Embryology_for_K12_Students|K12 students]] with a table comparing [[K12_Animal_Development_Times|Animal Development Times]]. | The table below lists the approximate development time for a large number of differen

Human embryonic stage 7 occurs during week 3 between 15 to 17 days. {{Carnegie stage 7 links}}

...e production a costly one, and we desire to express our obligations to the Carnegie Trust of the Scottish Universities for giving us a grant towards the expens ## [[1._An_Early_Ovum_imbedded_in_the_Decidua_(1908)#Table_of_Selected_Ova|Table of Selected Ova]]

[[File:Human Carnegie stage 1-23.jpg|400px|Human Embryo, Carnegie stages 1-23]] ...e human head at each stage of embryonic development starting at [[Carnegie stage 11]] at the beginning of Week 4. Making note of the relative head size, app

...56, 14 November 2019 (AEDT) Updated {{Hox}} page with human classification table. ...rt-stroke-vascular-diseases/congenital-heart-disease-in-australia/contents/table-of-contents CDK 14]. Canberra: AIHW.</ref>

==Stage 2== [[Carnegie stage 2]]

...jpg|thumb|alt=Mouse estrous cycle stage identification|Mouse estrous cycle stage identification{{#pmid:22514749|PMID22514749}}]] ...d descriptions are an excellent resource for learning how to determine the stage of the estrous cycle by visual observation or vaginal cytology."

File:Spiegel1626_table05.jpg|Table 5 Placenta Fetal and Maternal Side File:Spiegel1626_table05fig1.jpg|Table 5 Fig. 1 Placenta Fetal Side

Human embryonic stage 8 occurs during week 3 between 17 to 19 days. {{Carnegie stage 8 links}}

...ecular Development - Epigenetics|epigenetic]] changes at this early zygote stage.{{#pmid:27916276|PMID27916276}}{{#pmid:28818477|PMID28818477}} ...k 3) and is described as [[Carnegie stage 1|'''Carnegie stage 1''']]. This stage is followed by mitosis to form 2 blastomeres and then a solid cell mass cal

...embryo measured 11 mm CRL, probably corresponding to [[week 6]] [[Carnegie stage 16]]. A reconstructed model was also illustrated in the paper. '''Modern Pages:''' [[Carnegie stage 16]] | [[Week 6]]

...ersion]]. Added Carnegie collection [[Carnegie_stage_8#Carnegie_Collection|stage 8 images]]. ...32 Cell Stage‎‎|Quicktime version]] and [[Movie - Model Embryo to 32 Cell Stage‎‎|Flash version]].

...ttp://devblog.cloudsync.com/examples/wikiTableGenerator Generate Mediawiki table code]. ...e] may help. Also read the [http://meta.wikimedia.org/wiki/Help:Table Help:Table] information.

...]] added implantation animation, [[Morula]] and [[Blastocyst]] development stage images and references added to template pages. ...Development]] added [[:File:Human- Stage 22 integument 01.jpg|human embryo stage 22 skin image]] and movie of melanoblast migration in muse skin [[Quicktime

...to age fetuses based upon their bone ossification using embryos from the [[Carnegie Collection]]. ...t we find recognizable differences in size between male and female fetuses (table 2). Even with this reservation upon crown-rump length in later fetal life i

...image, just observe the anatomical structures present at this early fetal stage. ...10 week female fetus approximately 40 mm in size. This is showing an early stage of fetal development after the embryonic period (up to week 8).

[[File:Human Carnegie stage 10-23.jpg|thumb|300px|Carnegie Embryos]] ...collection numbering also incorporated the Blechschmidt embryo collection (Carnegie Nos. 10315-10434 ) in 1972, the collection embryos have now been returned t

{{Carnegie stage 1 links}} {{Carnegie stage 2 links}}

...have reported that they could distinguish carpal elements in the blastemal stage (Lewis, 1902; Schmidt-Ehrenberg, 1942; Haines, 1947). Chondrification begin ...ndrification, being cumulative, affords an excellent guide to the relative stage of development of embryonic limbs while the increase in the number of cente

* zygote ([[Carnegie stage 1]]) is the first diploid cell formed following fertilisation. :'''Links:''' [[Fertilization]] | [[Carnegie stage 1]]

...in Humans from week 3 to week 8 ({{GA}} week 5-10) and is divided into 23 Carnegie stages of embryonic development. There will also be a brief introduction to The [[#Carnegie_Stage_Table|Carnegie stages]] of the first 8 week of human development.

[[File:Human Stage14 neural02.jpg|thumb|Human Embryo CNS ([[Carnegie stage 14|stage 14]]) showing cranial nerve development]] {{Cranial Nerve Table}}

...assessment will be given in the practical class. Below in the collapsible table are examples of links from a virtual slide. There is also a [[Help:Virtual {{Virtual Slide Features - Stage 22 Liver}}

| [[File:Human embryo day 2.jpg|200px|Stage 2 Day 2]] | [[File:Human embryo day 3.jpg|200px|Stage 2 Day 3]]

Genetic code: Translation table 1 (Standard) Mitochondrial genetic code: Translation table 2 (Vertebrate Mitochondrial)

[[File:Stage14_sem2l.jpg|thumb|Embryo Stage 14]] ...and neck|1909 Head Malformations]] | [[Book - Contributions to Embryology Carnegie Institution No.68|1922 Aortic-Arch System]] | [[Paper - Evolutionary factor

...yo'''{{#pmid:22296782|PMID22296782}} "In all, 171 samples between Carnegie stage (CS) 17 and CS 23 were selected from MR image datasets of human embryos obt ...[[Carnegie_stage_14|stage 14]]) through to [[week 8]] ([[Carnegie_stage_23|stage 23]]) showing development of the [[A#auricular hillocks|auricular hillocks]

File:Stage10_SEM1.jpg|Stage {{CS10}} (22 - 23 days) File:Stage10_sem2.jpg|Stage {{CS10}} (22 - 23 days)

[[File:Stage11 sem81.jpg|thumb|Human face (Week 4, Stage 11)]] [[File:Stage14_sem2l.jpg|thumb|Human face (Week 5, Stage 14)]]

Human embryonic stage 6 occurs towards the end of week 2 at approximately 13-14 days. {{Carnegie stage 6 links}}

...y Models]] page with information about [[Embryology_Models#Carnegie_Models|Carnegie Models]]. ...o_Embryology_Carnegie_Institution_No.56|Contributions to Embryology Vol.12 Carnegie Institution No.56 (1921)]].

...Development_Animation_-_Week_1|Movie - Week 1 Cell Cleavage]] | [[Carnegie stage 1]] [[Carnegie stage 2]]

[[File:Heart Tube Segments.jpg|thumb|The embryo stage 10 heart tube]] ...C and magnetic resonance images from a study of 52 human fetuses (Carnegie stage 13-23). Time course of atrial, ventricular, and outflow septation were outl

{{Carnegie stage 11-14 image table}} [[Carnegie stage 15|'''Carnegie stage 15''']] <br>35 - 38 days

{{Animal icon table}} ...orm all the embryo, they are often described as the "germ layers" and this stage of development as the "gastrula". The same layers form the same structures

==Appendix 1 - Embryos In The Carnegie Collection== The Carnegie specimens of stages 2-23 are listed in the following tables.

...pment). Rat development is also generally 1 day behind that of mouse. (The table below gives details relating to the staging of rat development). ...albino rat (''Mus Norvegicus Albinus'') within the uterus from pronuclear stage to day 9.<ref name=Huber1915>{{Ref-Huber1915}}</ref>

[[File:Human Carnegie stage 10-23.jpg|thumb|Carnegie Embryos]] [[File:Mall Meyer1921 fig02.jpg|thumb|Carnegie Embryo key card]]

...stage 5|5]], [[Carnegie stage 6|6]], [[Carnegie stage 7|7]] and [[Carnegie stage 8|8]]). {{Monoygotic Twinning Table}}

| valign="bottom"|{{Stage 23 MRI movie 1}} These images of the Week 4 embryo (23 - 26 days, [[Carnegie stage 11|Stage 11]]) show the breakdown of the buccopharyngeal (oral) membrane.

{{Mouse Rat Pig table}} ...human pregnancy or to some earlier period? It would be possible to use the stage of development of a single organ such as the eye or heart to equate two mam

...|alt=Human embryonic cornea|Human embryonic cornea ([[Week 8]], [[Carnegie stage 22]])]] ...man embryonic cornea|Human embryonic cornea detail ([[Week 8]], [[Carnegie stage 22]])]]

| valign="bottom"|{{Neural stage 13 movie}} | valign="bottom"|{{Neural stage 22 movie}}

...em21.jpg|thumb|300px|Human embryo neural crest cells ([[Week 4]], Carnegie stage {{CS11}})]] ...ral02.jpg|thumb|300px|Human embryo neural ([[Week 5]], [[Carnegie stage 14|stage 14]])]]

The collection of embryos and fetuses of the Department of Embryology, Carnegie Institution of Washington, Baltimore, supplemented by specimens from the De Even in the initial stage of the gland’s development, the cells of the parathyroid are readily reco

[[Carnegie stage 14]] (CS14) orientation of the midgut loop and its mesentery during the 5th ...e midgut. Note the limited craniocaudal extension of the mesentery at this stage (10). The beige area identifies the region where the intestinal mesenchyme

{{Dobbin table}} ...to one of us (Hill) by Dr Roy Dobbin of Cairo. The embryo belongs
 to the stage characterised by the presence of a head-process with a well-developed chord

...etimes referred to as the Minot Collection, now forms part of the larger [[Carnegie Collection]]. The collection was described in detail by Minot (1905).<ref n [[Carnegie Collection]] - HDAC 7 Charles Sedgwick Minot Embryological Collection

...of the human embryo between Carnegie stage 19 to 23 in week 8 using the [[Carnegie Collection]] embryos. {{Carnegie stage 19 links}}

==Stage 23== {{Carnegie Collection stage 23 table}}

...in of the human embryo|Human Embryo vascular development (week 8, stage 20 Carnegie Embryo No. {{CE460}})]] ...embryo 50 mm. long (Carnegie Collection, No. 96)|Human embryo 50 mm long (Carnegie Collection, No. {{CE96}}.]]

...the [[Harvard Collection|Harvard Embryological Collection (H.E.C.)]] and [[Carnegie Collection]]. ...were studied; of those between 3.0 and 6.6 mm., forty-four series in the [[Carnegie Collection]]. The series in the Harvard Collection which best displayed the

...44 paper by Davies describes early human development in week 2 ([[Carnegie stage 5]]). {{Carnegie stage 5 links}}

...Mateer Embryo" was later catalogued as Carnegie Embryo {{CE1399}} Carnegie stage {{CS8)). '''Modern Notes:''' Carnegie stage {{CS8)) | Embryo {{CE1399}} | [[Week 2]] | [[Week 3]]

Stage 16 to 18 Stage 15 to 22

...b|link=External Genital Male Development Movie|[[Week 9]], the indifferent stage of external genitalia.]] ...nds into the glans penis and clitoris, respectively, during the ambisexual stage (9 weeks) and thus appears to be an androgen-independent event. (2) The ent

* Carnegie stage {{CS19}} - Thick stalk with remnant of lumen (Rathke’s pouch); angiogenes * Carnegie stage {{CS20}} - Long, slender stalk.

...the definitive GIT tube is formed and then at the overview of the Carnegie stage {{CS22}} embryo GIT from one end to the other. The tract now has a differen ...comparing the GIT appearance with the earlier embryonic {{CS13}} Carnegie stage.

...Bardeen]] describes intestinal development using human embryos from the [[Carnegie Collection]] and Wisconsin Collection. ...junum and coils 4, 5 and 6, the ileum we find from the data given by Mall (table 1, footnotes) that the ileum is about four times as long as the jejunum dur

| valign="bottom"|{{Embryo stage movie 1}} ...way again by clicking "hide". A very useful tool, don't forget to give the table a name that hints at its content.

[[File:Stage11 sem100.jpg|thumb|300px|Human embryo ([[week 4]], [[Carnegie stage 11]]) Somites]] ...at regular time intervals. The process is sequential and therefore used to stage the age of many different species embryos based upon the number visible som

[[:File:Human_Carnegie_stage_1-23.jpg|Carnegie stage 1-23]] [[File:Stage14 sem2b-limb.jpg|thumb|Human Embryo stage 14 SEM]]

...ge_22_image_217.jpg|thumb|300px|Cerebrum development human embryo (week 8, Stage 22)]] | valign="bottom"|{{Neural stage 13 movie}}

..._22_image_085.jpg|thumb|300px|Developing Human Spleen ([[Carnegie stage 22|stage 22]])]] ...ing the human embryonic period'''{{#pmid:25403423|PMID25403423}} "Between Carnegie stages (CSs) {{CS14}} and {{CS17}}, the spleen was usually recognized as a

Carnegie Stage 4 (stages 1-23 describing key steps in embryonic development) represents th {{Monoygotic Twinning Table}}

==Stage 7== [[Carnegie stage 7]]

Published by the Carnegie Institution of Washington Washington, 1920 The body could be safely straightened for crown-rump measurement down to the stage of 35 or 40 mm. long. Specimens smaller than this were measured without dis

{{Eye Timeline table}} * Carnegie stage {{CS19}} - Optic nerve small, slender. Lumen practically whole length of st

Theiler Stage 1 - Fertilization Theiler Stage 2 - Dividing egg stage 2-4 cells. Zona pellucida present. First cleavage occurs at about 24 hours.

...resonance imaging. The cartilaginous femur was first observed at Carnegie stage 18. Major anatomical landmarks were formed prior to the initiation of ossif ...and Somite, Vertebral Column, Limb in combination with serial sections and Carnegie images.

...portion of the pancreas. The study include embryos and fetuses from the [[Carnegie Collection]]. ...be shown, to trace the relation of an island to its acinus. In this early stage it was not possible to find any arrangement of cells suggestive of the for

{{Neural Table}} ...13 image 098.jpg|thumb|300px|Brain primary vesicle development ([[Carnegie stage 13]])]]

...{{CE1584}} [[Carnegie stage 19|stage 19]]; {{CE1535}} [[Carnegie stage 23|stage 23]]. '''Modern Notes:''' {{embryonic}} | {{CRL}} | [[Carnegie Collection]]

...s described initially as [[Carnegie stage 3|'''Carnegie stage 3''']]. This stage is followed by blastocyst hatching and implantation. ..., and ERK pathways efficiently (>70%) form blastoids generating blastocyst-stage analogs of the 3 founding lineages (>97% trophectoderm, {{epiblast}}, and p

...paper describes an early human embryo (Pfannenstiel III) about [[Carnegie stage 11]] occurring in [[week 4]]. From the Collection of Prof. Pfannenstiel, Gi | {{Human embryo Pfannenstiel III table}}

...eural groove closing to neural tube, early week 4 <br>([[Carnegie_stage_10|Stage 10]])]] <html5media height="320" width="300">File:Stage 16 MRI 3D03.mp4</html5media>

...clavicle contains a relatively large amount of cartilage at a fairly early stage. In man the cartilage is of a peculiar kind called by Mall a ‘precartilag ...nnection, but were separated by the investing perichondrium. In the 19-mm. stage (crown-rump measurement) a bony bridge develops and connects the two center

...cavity, it is not readily visible in the external views of the embryonic (Carnegie) stages of development. The tongue has {{neural crest}} origins and muscle ...ape and size, {{Hippo}} signalling and cell proliferation in a region- and stage-specific manner'''{{#pmid:34697858|PMID34697858}} "The mammalian tongue dev

...Mckay and co-authors describes human [[Carnegie stage 14|Carnegie horizon (stage) 14]] embryos. Currently only a brief abstract is included on this page. '''Modern Pages:''' [[Carnegie stage 14]] | [[Week 5]] | [[Embryology History - Arthur Hertig|Arthur Hertig]]

[[File:Stage22_mesonephros.jpg|thumb|Male urogenital development (stage 22)]] ...ent of the human female reproductive tract is reviewed from the ambisexual stage to advanced development of the uterine tube, uterine corpus, uterine cervix

==Stage 10== [[Carnegie stage 10]]

...appearance of the upper and lower limb buds are more like a later Carnegie stage {{CS16}} embryo occurring in week 6, 37 - 42 days, CRL 8 - 11 mm. This stage in the growth of the brain is particularly interesting in that it represent

...duct. ...Descent of the thyroid gland was not evident after the CRL 20 mm stage (6 weeks): the gland appeared to retain its position at the level of the th | [[File:Stage13 and 22 thyroid development a.jpg|600px|Stage 13 and Stage 22 thyroid development]]

...'{{#pmid:26995337|PMID26995337}} "We selected 37 human embryos at Carnegie Stage (CS) {{CS11}}-{{CS13}} (28-33 days after fertilization) and three-dimension |+ '''Late Embryonic Duodenum''' (Week 8, {{GA}} week 10, Stage {{CS22}})

...genesis of the thyroid follicles (Norris, ’16), has been carried on at the Carnegie Institute of Embryology and at the University of Minnesota under the superv This study is based upon the collection of human embryos in the Carnegie Institute of Embryology at Baltimore and upon those in the Anatomical Labor

Use the links to [[Carnegie stage 7]], [[Carnegie stage 8]] and [[Carnegie stage 9]] to see a number of different views of the human embryo in the third wee ==Stage 7==

| valign="bottom"|{{Embryo stage movie 1}} His's Normentafel (Normal Table)

...ed developmental periods, including all periods between the gonadal tissue stage and foetal testes development. The number of NANOG, DDX4, DAZL, DPPA3 and 5 ...quently cryopreserved by a vitrification method. When the 4-cell to morula stage of cryopreserved embryos were non-surgically transferred into the uteri of

[[File:Carnegie stage 13 caudal trunk.jpg|thumb|Human Embryo (Carnegie stage 13) caudal trunk<ref><pubmed>18689800</pubmed>| [http://hmg.oxfordjournals. ...arches] | [http://www.ncbi.nlm.nih.gov/books/bv.fcgi?&rid=dbio.table.3135 Table 13.2. Some derivatives of the pharyngeal arches]

[[File:Stage 13 kidney sections.jpg|right]] ...re_-_Endocrine_Development#Endocrinology_-_An_Integrated_Approach|Detailed Table of Contents]] | [http://www.ncbi.nlm.nih.gov/books/NBK22 Bookshelf Link]

* Embryo Bs - 26-27 pairs of somites [[Carnegie stage 12]] in [[Week 4]] (Somite pairs 21 - 29) * Embryo Fu - 28 pairs of somites [[Carnegie stage 12]] in [[Week 4]] (Somite pairs 21 - 29)

By A. M. Hain (Carnegie Research Fellow), Institute of Animal Genetics, University of Edinburgh, ...were largely defrayed by grants from the Medical Research Council and the Carnegie Trust for the Universities of Scotland.

{{Carnegie Collection stage 20 table}} {{Carnegie numbered embryo links}}

| valign="bottom"|{{Stage 23 MRI movie 1}} ...lt=Human embryo Stage 13 oral cavity floor|Human embryo pharyngeal arches (Stage 13 oral cavity floor)]]

Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland, and Department of Zoology, ...Mann states (p. 254) that “the eye muscles in man seem, at their earliest stage, to be represented by a single undifferentiated mass. The earliest trace of

| [[File:Stage5 bf11L.jpg|alt=Human embryo Carnegie stage 5|400px]] * '''conceptus''' completes implantation during this stage.

...muno-histological overview of the fetal human middle ear during a critical stage of tympanic cavitation was provided. A critical analysis of previously repo ...les in the middle ear were independent in different locations. At Carnegie Stage 17 a homogeneous interzone clearly defined the incus and malleus anlagen. T

...ibes gastrointestinal tract smooth muscle development using a number of [[Carnegie Collection]] embryos. {{Carnegie Collection fetal table}}

...embryo spina bifida. This was an early publication in the series from the Carnegie Institution of Washington called [[Book_-_Contributions_to_Embryology|Contr [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]]

...by the Department of Embryology, sincere appreciation is expressed to the Carnegie Institution of Washington, and to Dr. G. W. Corner, Director. The author is ===Table 1 - Human Embryos of the Carnegie Collection used in this Study===

...f brain function. The fetal to neonatal period is well known as a critical stage in brain development. In this study, we evaluate the network topography of {{Human fetal length-weight table}}

...Mckay and co-authors describes human [[Carnegie stage 13|Carnegie horizon (stage) 13]] embryos. Currently only a brief abstract is included on this page. '''Links:''' [[Carnegie stage 13]] | [[Week 5]] | [[Embryology History - Arthur Hertig|Arthur Hertig]]

** ventral bud later ([[Carnegie stage 13|stage 13]] - [[Carnegie stage 14|stage 14]]) During week 6 about day 41 ([[Carnegie stage 17]], {{GA}} week 8) the stomach rotation brings the smaller ventral pancre

...nell University.‘ Sixteen of the human series are in the collection of the Carnegie Institution of Embryology,” and for the privilege of studying these embry ...thickened throughout the neural plate. As the neural folds form (1-somite stage, fig. 1 A; see also Adelmann, ’25) mesenchyme cells fill the cavity of

...a mystery because of the difficulties in accessing primate embryos at this stage. Here, we report the establishment of an in vitro culture (IVC) system that ...oplasmic sperm injection (ICSI), preimplantation development to blastocyst stage, and embryo transfer techniques are described. With baboon ART methodologie

...n numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

...omandibular joint, that connecting the mandible to the skull. Pater uses [[Carnegie Collection]] embryos: {{CE1318}}, {{CE1455}}, {{CE3990}}, {{CE5652}}, {{CE6 ...The investigation was supported by a grant from the fellowship fund of the Carnegie Institution of Washington and by the Department of Anatomy, Medical College

Department of Embryology, Carnegie Institution of Washington ...d was in an embryo (No. {{CE234a}}) 80 mm. crown-rump length. The youngest stage was an embryo (No. {{CE588}}) 4 mm. greatest length, at which time the oti

==Stage 5== [[Carnegie stage 5]]

[[File:Stage22_mesonephros.jpg|thumb|Male urogenital development (stage 22)]] ...nt study provides new insights into the function of Bmp signaling at early stage for the initiation of GT outgrowth." {{BMP}}

(From the Department of Genetics, Carnegie Institution of Washington, Cold Spring Harbor.) ...rt with the beginning of the embryo proper and not include the preliminary stage or the extraembryonic tissues. This proposition is shown to be justified b

His's Normentafel (Normal Table) [[Carnegie Collection]]

[[File:Stage 22 image 084.jpg|thumb|Human Liver (week 8, GA week 10)]] ...ween E11.5 and E14.5, but their maturation states at a given developmental stage were heterogeneous. Even more surprising, the number of proliferating cells

...egie Staging:''' A 27 somite stage embryo would be similar to a [[Carnegie stage 12]] (26 - 30 days), caudal neuropore closes, Somite Number 21-29. '''Modern Notes:''' [[Carnegie stage 12]] | [[Week 4]] | {{neural}} | {{somite}}

To eliminate a too frequent use of 'oestrum' in its various forms, the stage will be designated as shown below: IMo J condition. Stage D. In the mouse during the dioestrous interval the vulva is very inconspicu

...n numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

...sinus. These human embryos are [[Carnegie Embryos]] and fetuses from the [[Carnegie Collection]]. In the adult, the endolymphatic sac regulates the inner ear Department of Embryology, Carnegie Institution of Washington, Johns Hopkins Medical School, Baltimore, Marylan

...groove closing to neural tube<br>Embryo early week 4 ([[Carnegie_stage_10|Stage 10]])]] ...e section week 8|Spinal cord transverse section<br>Embryo week 8 (Carnegie Stage {{CS22}})]]

in the stage 13/14 embryo (shown below) the otic placode has sunk from the surface ectod Primary palate, fusion in the human embryo between stage 17 and 18, from an epithelial seam to the mesenchymal bridge.

...mbryo 841|841]], [[:Category:Carnegie Embryo 5537|5537]], and [[:Category:Carnegie Embryo 6521|6521]]). Note Mall's reference is cited as 1913, but it is the {{Heart - Historic References table}}

===Table A. Number of females per 1,000 males. (From Rauber.)=== ===Table B. Changes in sex-ratio with advancing age. (From Prinzing.)===

{{Carnegie No.59 Header}} ...of its purpose, has placed at my disposal the collection of embryos of the Carnegie Laboratory of Embryology, together with models, notes, and such other relat

...les R. Essick describes human embryonic {{neural}} development using the [[Carnegie Collection]] embryos. [[Carnegie Embryos]] used in this study: {{CE227}}, {{CE75}} {{CE86}} {{CE145}} {{CE1

...e_22_image_055.jpg|thumb|Human embryonic hypothalamus ([[Carnegie_stage_22|Stage 22]])]] ...n region]], shown by [[O#optic stalk|optic stalk]]<br>([[Carnegie_stage_13|Stage 13]])

[Embryo No. {{CE460}} is classified as a [[Carnegie stage 20]] (week 8, {{GA}} week 10) embryo. The following pages also relate to th ...he conditions found in [[:Category:Carnegie Embryo 460|embryo No. 460]] ([[Carnegie Collection]]), which are shown in figures 7 and 8, to the theory that "onto

[[File:Stage10_sem9.jpg|thumb|Stage 10 neural groove to tube]] ...opment of the [[N#neural plate|neural plate]] region at the embryonic disc stage.

...olman numbers the embryos examined in sequence 1-30, but also gives Mall's Carnegie embryo numbers in the tabulation. Carnegie Embryo {{CE186}}

|[[File:Mark_Hill.jpg|50px|left]] This historic 1956 paper describes using [[Carnegie Collection]] early human development in week 2 and 3. [[Carnegie Embryos|Carnegie Embryos in this paper]]: {{CE8698}} | {{CE8794}} | {{CE8663|}} | {{CE8663}

...age 22 vomeronasal organ.jpg|thumb|300px|Human embryo ([[Carnegie stage 22|stage 22]]) nasal epithelium development.]] Carnegie stage {{CS13}}

{{Carnegie stages}} [[Carnegie stage 7]]

...are not Carnegie stages, use the embryo CRL to approximately convert to [[Carnegie Stages]]. ...bryology_15|Historic - Urogenital Development]] | [[Carnegie Embryos]] | [[Carnegie Collection]]

{{Carnegie stages}} [[Carnegie stage 7]]

..._.28Post_Fertilization.29_of_Susceptibility_to_a_Drug-Induced_Malformation|Table 2]]. ...ent_and_maldevelopment#Table_1._Comparison_of_Human_and_Murine_Development|Table 1]]).

| '''Human Embryo''' (8 weeks, [[Carnegie stage 22|stage 22]]) adrenal gland showing the fetal and permanent adrenal cortex. Note th Table modified from review.{{#pmid:14594453|PMID14594453}}

...re_-_Endocrine_Development#Endocrinology_-_An_Integrated_Approach|Detailed Table of Contents]] | [http://www.ncbi.nlm.nih.gov/books/NBK22 Bookshelf Link] [[File:Stage22 mesonephros.jpg|thumb|Stage 22 mesonephros]]

...4 neural crest cells|Human embryo neural crest cells ([[Week 4]], Carnegie stage {{CS11}})]] ...migration occurs at different rates along the embryo axis between Carnegie stage 11 to 13 in week 4.

...em, organ, tissue or region can be followed in older embryos from stage to stage. The various embryos must be classified and standardized, the membranes mus ...ay turn out that the organs are developed unequally in embryos of the same stage and this would be an observation of great importance.

By A. M. Hain (Carnegie Research Fellow), The Institute of Animal Genetics, Edinburgh University, ...y defrayed by grants (to A.M.H.) from the Medical Research Council and the Carnegie Trust for the Universities of Scotland.

...paper by Watson described fetal human seminal vesicle development using [[Carnegie Collection]] specimens. James Buchanan Brady Urological Institute, Johns Hopkins Hospital, and the Carnegie Laboratory of Embryology, Baltimore, Maryland

...he human were lacking except for some modeled specimens at the four somite stage (Heuser and Corner, 12) which show a shallow foregut. The sides of the head grooves connecting into it. At the seven somite stage of human embryos (Dandy, 5) a single pair of pharyngeal pouches in the form

...ceptus fully implanted|Human conceptus fully implanted ([[Carnegie_stage_5|Stage 5]]).]] * '''Dynamics of {{trophoblast}} differentiation in peri-{{implantation}}-stage human embryos'''{{#pmid:31636193|PMID31636193}} "Single-cell RNA sequencing

==Stage 12== [[Carnegie stage 12]]

...clearly concerned with atretic follicles. Also in a majority of cases the stage of the menstrual cycle at which the ovary was removed was not known. Some k * 2 In press: Carnegie Publications, Contributions to Embryology.

* '''Contributions to Embryology''' - [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington Series]] A historic series of papers published by the Carnegie Institution of Washington early in the 20th Century.

...result of an invagination of the coelomic epithelium - [[Carnegie_stage_18|stage 18]] ...''' - duct grows independently from the invagination - [[Carnegie_stage_19|stage 19]]

...clearly concerned with atretic follicles. Also in a majority of cases the stage of the menstrual cycle at which the ovary was removed was not known. Some k * 2 In press: Carnegie Publications, Contributions to Embryology.

Carnegie Institution of Washington, Department of Embryology ...detail in the near future in Contributions to Embryology, published by the Carnegie Institution of Washington.

[[File:Human_idiogram.jpg|200px]] [[File:Stage22 mesonephros.jpg|300px|Stage 22 mesonephros]] [[File:Historic-testis.jpg|120px]] [[File:Historic-ovary.j [[File:Stage22 mesonephros.jpg|thumb|Stage 22 mesonephros]]

...different species. Note that this online version is only at an early draft stage of editing. Department of Embryology, Carnegie Institution of Washington, Baltimore, Mn.

...and Somite, Vertebral Column, Limb in combination with serial sections and Carnegie images. | [[Carnegie stage 13]]

...the outer border of the common carotid artery and the thyroid. After this stage the more rapid growth of the neck of the embryo than of thymus III and the ...the glands in the different embryos and foetuses is given in the following table.

...pment of the trachea and esophagus and includes several embryos from the [[Carnegie Collection]]. Department Of Embryology, Carnegie Institution Of Washington, Baltimore, Maryland

...evelopment of pregastrulating embryos (E5.5) until the hind limb formation stage (E11). Late gastrulating embryos (E7.5) are grown in rotating bottles in th {{Mouse Rat Pig table}}

...evelopment of the limbs, body-wall and back using human embryos from the [[Carnegie Collection]]: {{CE2}}, {{CE12}}, {{CE22}}, {{CE43}}, {{CE76}}, {{CE106}} {{ In the accompanying table a list is given of the embryos utilised those marked with an asterisk have

...pment. This paper includes descriptions of the venous system of several [[Carnegie Collection]] human embryos as well as human embryos from the [[Harvard Coll ...[[Carnegie stage 17]]) | [[:Category:Carnegie Embryo 841|841]] ([[Carnegie stage 18]])

# Week 4 Embryo - Stage 13 ([[Carnegie_stage_13_-_serial_sections|2012]]) # Week 8 Embryo - Stage 22 ([[Carnegie_stage_22_-_serial_sections|2012]])

[[File:Stage18 em11.jpg|thumb|200px|Human Embryo Face ([[Week 7]], Carnegie stage {{CS18}}, 44 - 48 days, CRL 13 - 17 mm)]] ...e.jpg|thumb|200px|Human Embryo clefting. (Week 6, {{GA}} week 8, Carnegie stage {{CS16}}, ventral view)]]

...o the method of Born. A list of these embryos is given in the accompanying table : Table op Embryos Modeled. Lengtb in mm.

...p://embryology.med.unsw.edu.au/wwwhuman/Stages/Images/stage14compare23.jpg Stage Comparison] ...e great change beween the middle of embryological development to the final stage.

University of Chicago Embryo H279 was added to the Carnegie Collection as Embryo {{CE3709}}. ...t of Anatomy, The University of Chicago, and the Laboratory of Embryology, Carnegie Institution of Washington

...and others on human vertebral column development using embryos from the [[Carnegie Collection]].{{#pmid:517765|PMID517765}}{{#pmid:7216919|PMID7216919}}{{#pmi {{Axial Skeleton table}}

{{Carnegie stage 11-14 image table}} [[Carnegie stage 15|'''Carnegie stage 15''']] <br>35 - 38 days

File:Gray0178.jpg|Embryo 24 mm (outer aspect, about [[Carnegie stage 22]]) File:Gray0179.jpg|Embryo 24 mm (inner aspect, about [[Carnegie stage 22]])

Cultured eight-cell morulae and four-cell-stage mouse embryos to the Blastocyst sage. ...of successful attempts to culture mouse embryos in vitro to the Blastocyst stage (Nagy et.al. 2003).

...evelopment of the limbs, body-wall and back using human embryos from the [[Carnegie Collection]]: {{CE2}}, {{CE12}}, {{CE22}}, {{CE43}}, {{CE76}}, {{CE106}} {{ In the accompanying table a list is given of the embryos utilised those marked with an asterisk have

At this stage the zona pellucida may still be adherent to the trophoblast. The total numb ...see [[Template:Macaque Placental Timeline table|Macaque Placental Timeline table]] from this data.

...he evaluation of a range of measures in the kidneys, such as developmental stage, rate and success, injury, and disease processes, relies on obtaining infor ...ormed during human embryogenesis. Fifty human embryos between the Carnegie stage {{CS14}} and {{CS23}} were selected from the [[Kyoto Collection]] at the Co

...s human embryo collection that will eventually form the beginning of the [[Carnegie Collection]]. See also later paper by Streeter, GL. [[Book - Contributions to Embryology Carnegie Institution No.24|The Developmental Alterations in the Vascular System of t

...rain vascular development includes descriptions of many embryos from the [[Carnegie Collection]] including: {{CE84}}, {{CE96}}, {{CE144}}, {{CE199}}, {{CE234a} ...age of other structures which have not yet made their appearance. For each stage it is an efficient and complete going-mechanism, apparently uninfluenced by

...gie_stage_15|stage 15 embryo]] (28 - 32 days) than the [[Carnegie_stage_12|stage 12 embryo]] (26 - 36 days). ...as Carnegie No. {{CE2}} and [[Carnegie_stage_15|stage 15 embryo]] in the [[Carnegie Collection]].

{{Shaw table}} ...esis dominates and angiogenesis appears to be secondary. This intermediate stage bridges over a gap in the history of our knowledge of the development of th

...ory_tract.jpg|thumb|300px|Respiratory system overview ([[Carnegie_stage_13|stage 13]])]] ...pmid:37602519|PMID37602519}} "Imaging data of 41 human embryo specimens at Carnegie stages (CS) 16-23 (equivalent to 6-8 weeks after fertilization) belonging t

[[File:Stage14 sem2cl.jpg|300px|thumb|right| '''Figure 4.''' Stage 14]] ==Carnegie Stages==

For scanning electron micrographs of the Carnegie stages of the early human embryo from embryos collected by Dr. Vekemans and For providing images and stage information on the embryonic development of the bat. Both researchers are i

...ment. The paper uses descriptions of several of the early embryos of the [[Carnegie Collection]]: {{CE22}}, {{CE43}}, {{CE109}}, {{CE143}} and {{CE164}}. Also * Embryo No. {{CE22}} - This embryo was classified as [[Carnegie stage 21|Stage 21]] in [[Week 8]].

[[Image:CSt3.jpg|thumb|Human Blastocyst (Carnegie Stage 3)]] ...pment, the embryonic stem cell with totipotential occurs at the blastocyst stage, mainly in the first and second week of development. After this period the

...[[:Category:Carnegie Embryo 2|Carnegie Embryo No. 2]] [[Carnegie_stage_15|stage 15]] was first described in {{Ref-Mall1891}} {{Carnegie stage 15 links}}

{{Neural Table}} [[File:Stage10_sem6.jpg|thumb|Human embryo neural groove ([[Carnegie stage 10]])]]

{{Carnegie stage 19 links}} {{Carnegie stage 20 links}}

</ref> The dimensions and approximate ages of these embryos are given in the table on the following page. ===Table of Young Human Ova===

...hose gonads, if present at all, are in the indifferent stage, of which the table shows ten; the second group includes the females, and the third, the males. ===Table 1===

...y projection or camera luclda as whole embryos and subsequently sectioned (Table 1). ===Five-Somite Stage===

{{Seminiferous Tubule puberty table}} ...e excessive expression of transposable elements. Upon entry to the meiotic stage, XYsry- oocytes demonstrated extensive defects, including the impairment of

...yos are derived from Grosser (1924), those of the older specimens from the table of Mall (1910) according to the crown-rump measurement in embryos and foetu ...g their development, erythroblasts multiply by mitotic division. In a late stage of development they probably do not multiply and those with pyknotic nuclei