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...re are also sets of [[Carnegie stage 22 - selected serial sections]] and [[Carnegie stage 13 - serial sections]]. | [[:File:Stage 22 image 093.jpg|93]]

[[File:Human embryo day 2.jpg|thumb|300px|Human morula (day 2){{#pmid:19924284|PMID19924284}}]] A key event prior to morula formation is "compaction", where the 8 cell embryo undergoes changes in cell morphology and cell-cell adhesion that initiates

...hilly 1987|link=Embryology History - Ronan O'Rahilly|Ronan O'Rahilly (1987 Carnegie Labs)]] ...t study,<ref name=Weller1933>{{Ref-Weller1933}}</ref> used the following [[Carnegie Collection]] embryos: stage {{CS9}} (No. {{CE1878}}), {{CS10}} ({{CE391}};

...in the embryo as at birth, but the deformities of the head and neck of the embryo are of such a nature that it can not survive long enough to admit of compar ...xomphaly. Other anomalies, however, are more difficult to recognize in the embryo as sharply defined malformations.

...23 paper by Finley describes vascular development in the head of the human embryo. ...day 50). Its development and position was historically used to stage late Carnegie embryos.

...23 paper by Finley describes vascular development in the head of the human embryo. ...day 50). Its development and position was historically used to stage late Carnegie embryos.

...Stage11 sem21.jpg|thumb|300px|Human embryo neural crest cells ([[Week 4]], Carnegie stage {{CS11}})]] ...:Human Stage14 neural02.jpg|thumb|300px|Human embryo neural ([[Week 5]], [[Carnegie stage 14|stage 14]])]]

[[Carnegie Collection]]: {{CE45}}, {{CE75}}, {{CE86}}, {{CE95}} ...en Kautenhirns''"; a description of these is accordingly unnecessary. Each embryo has been studied in serial sections and from tliese sections a few, at diff

[[File:Human Carnegie stage 10-23.jpg|thumb|300px|Carnegie Embryos]] ...collection numbering also incorporated the Blechschmidt embryo collection (Carnegie Nos. 10315-10434 ) in 1972, the collection embryos have now been returned t

...}</ref> Later in 1921 along with Mall published a review of abnormal human embryo development.<ref>{{Ref-Mall1921}}</ref> ...lips of the blastopore (in the late gastrula stage) to other parts of the embryo and found that as expected they differentiated into structures characterist

...bers the embryos examined in sequence 1-30, but also gives Mall's Carnegie embryo numbers in the tabulation. Carnegie Embryo {{CE186}}

...many different locations and differentiate into many cell types within the embryo. This means that many different systems (neural, skin, teeth, head, face, [[File:Carnegie stage 13 caudal trunk.jpg|thumb|Human Embryo (Carnegie stage 13) caudal trunk<ref><pubmed>18689800</pubmed>| [http://hmg.oxfordjou

...of true embryonic stem cells capable of forming all cell types within the embryo. In mammals, the {{trophectoderm}} will form key cells ({{trophoblast}}) of ...({{GA}} week 3 and 4) and is described initially as [[Carnegie stage 3|'''Carnegie stage 3''']]. This stage is followed by blastocyst hatching and implantatio

...function of long noncoding RNAs (lncRNAs) and small RNAs during oocyte-to-embryo transition in mammals. LncRNAs are an assorted rapidly evolving collection ...miR-183) are differentially expressed in the CVG compared to NC and OV at Carnegie developmental stage {{CS13}}. We further identified transcription factors t

Development of this system commences in the embryo, continues through the fetal period then with key changes around birth, onl ...Reproductive development has a long maturation timecourse, begining in the embryo and finishing in puberty. (More? [[Puberty Development]])

...uld be clearly timed in the mouse and found in the literature on the human embryo. ...lation and fertilization times were unascertainable so that the age of the embryo is determined by the mating time plus or minus 30 minutes. The time for mat

...e 19|stage 19]]; {{CE1584}} [[Carnegie stage 19|stage 19]]; {{CE1535}} [[Carnegie stage 23|stage 23]]. '''Modern Notes:''' {{embryonic}} | {{CRL}} | [[Carnegie Collection]]

..., the embryo possesses a number of relatively large blood vessels. Thoma ('93) mentions that the paired dorsal aortae are present, and in a schematic dra ...s with the venous end of the heart and with the entire dorsal aorta of the embryo opposite the zone of the myotomes." That the heartbeat has much to do with

! width=120px|Carnegie Stage File:Keith1902 fig093.jpg|Fig. 93. Section showing the Uro-genital Sinus in the male foetus.

...:Stage10_neural_sm.jpg|thumb|300px|Neural groove closing to neural tube<br>Embryo early week 4 ([[Carnegie_stage_10|Stage 10]])]] ...transverse section week 8|Spinal cord transverse section<br>Embryo week 8 (Carnegie Stage {{CS22}})]]

...w that the malleus and incus are differentiated in the human and mammalian embryo. (1) The otocyst — an area or plaque of ectoderm covering the head of the embryo above the first visceral cleft which becomes invaginated in a saccular

{{Carnegie stage 19 links}} {{Carnegie stage 20 links}}

...t "hatching" from the zona pellucida ([[Week 1|week 1]] [[Carnegie_stage_3|Carnegie stage 3]])]] ...hese structures is needed to elucidate their potential role in early human embryo development."

...e free to use our judgment in methods of fixation and preservation. If the embryo is perfectly fresh or possibly living, we use, of course, the most refined ...straight and other measurements and weights also are taken. The age of the embryo is estimated on the basis of weight, crown-rump, and foot length, and the e

...bryo 391|Carnegie No. 391]] | [[Week 4]] | [[Somitogenesis|8 somites]] | [[Carnegie Collection]] [[Book - Contributions to Embryology|Contributions to Embryolo ...n. Wash. Publ. 362, Contrib. Embryol, 17, 1-67). Plaster models now at the Carnegie laboratory were made by [[Embryology History - Osborne Heard|O. Heard]] und

...of the lower axial skeleton and lower limbs using human embryos from the [[Carnegie Collection]]: {{CE2}}, {{CE22}}, {{CE45}}, {{CE62}}, {{CE109}}, {{CE144}}, [[Embryology History - Charles Bardeen|Charles Bardeen]] | [[Carnegie Embryos]]

...Mall describes the human embryos in the collection that would become the [[Carnegie Collection]]. There is also a [[:File:1904 - Catalogue of the collection of [[Carnegie Collection]] | [[Carnegie Embryos]]

...rain vascular development includes descriptions of many embryos from the [[Carnegie Collection]] including: {{CE84}}, {{CE96}}, {{CE144}}, {{CE199}}, {{CE234a} ...Developmental Alterations in the Vascular System of the Brain of the Human Embryo=

...indicate the absence of pouches. Weller (30) described a two somite human embryo which according to his description possessed the first pharyngeal pouch. Th Corner (4) described the foregut of a 10-somite human embryo, as being compressed dorso-ventrally with the anterior end immediately unde

...y Atlas of the 13-mm. Pig Embryo. (Prefaced by younger stages of the chick embryo.) The Wistar Institute Press, Philadelphia, iv & 104 pp. ...ggregation and phagocytosis of embryonic cells. Jour. Morph., Vol. 80, pp. 93-111.

...essum.' For Driesch ('93) has demonstrated for the sea-urchin and Wilson ('93) for Branchiostoma that the development of various 'monozygotic'<ref>This v ...one-third of the egg seems to be necessary for the development of a whole embryo.

...pigenesis.”” On theoretical grounds, he held that the various parts of the embryo were contained in the egg and became visible as they increased in size. The ...e more general characteristics of any large group of animals appear in the embryo earlier than the more special characteristics.

...ltttea associated with 7‘/3- and 9‘/3-day normal pregnancies respectively (Carnegie nos. {{CE8020}}, {{CE8215}}). Unfortunately, these sections, which were sta ===A 12- to 13-Day Pregnancy, Carnegie No. {{CE8558}}, S46-2767===

...l the cytoplasm is divided into two giving nse to a two cell st ige of the embryo or ovum (Fig 28B and C) Each of the daughter cells contains an equal number ...stage stage F —eight cell stage ( \fter Lev is and Hartman 1933 ) of tf e Carnegie Institution of Washington y c *00

The measurements of the embryo are as follows: C.R., crown-rump or sitting height; C.H., crown—hee1 or s <div id="Carnegie Embryo 6"></div>

Department of Embryology, Carnegie Institution of Washington, Baltimore, Mn. ...th its associated tiny mass of protoplasm, which alone will make the chick embryo, becomes all but lost on the surface of the yolk. But that speck of protopl

....jpg|90px|left]] This 1940 paper by Keith describes abnormalities in human embryo development. ...ge L. Streeter,‘ who has presided over the Department of Embryology of the Carnegie Institution, Washington, for many years and whose important contributions t

...e show that although blood cell formation in its early stages in the human embryo and especially the yolk sac follows the same general lines as in other mamm ...y with the magma strands of the chorionic cavity. At the caudal end of the embryo these yolk sac mesodermal cells, which will give rise to the yolk sac mesen

'''Brief timeline of rabbit embryo model use''' *1890- Walter Heape succeeded first mammalian embryo transfer

...pment of the trachea and esophagus and includes several embryos from the [[Carnegie Collection]]. Department Of Embryology, Carnegie Institution Of Washington, Baltimore, Maryland

....jpg|90px|left]] This historic 1941 paper by Gilmour describes early human embryo blood formation. ....065 x 0.045 mm. Age about 16 days, probably slightly younger than Peters’ embryo (1899).

...pmid:37602519|PMID37602519}} "Imaging data of 41 human embryo specimens at Carnegie stages (CS) 16-23 (equivalent to 6-8 weeks after fertilization) belonging t ...ng to 68% of genes that are down-regulated upon Nkx2-1 deletion, including 93% of known AT1 genes, but near-background binding to up-regulated genes. Our

...description of the transitory cavities in the corpus striatum of the human embryo. =Transitory Cavities in the Corpus Striatum of the Human Embryo=

.... Parthenogenese spontanee dans I'ovaire du cobaye. Bull. Soc. Anat. Paris 93, 724. ...pregnant women and by freshly implanted anterior lobe. J. Am. Med. Assoc. 93, 276.

...foetus. Edinburgh. , 1904. Manual of antenatal pathology and hygiene. The embryo. Edinburgh. DANDY, W. E., 1910. A human embryo with seven pairs of somites measuring about 2 mm. in length. Amer. Jour. An

=Chapter III. The Human Embryo= ==Calculation of the Age of the Human Embryo==

...bryology_History_-_Wilhelm_His|Wilhelm His]], but didn't receive the first Carnegie specimen until his position at Johns Hopkins University. ...a poor way to organize embryos. It is very difficult to accurately age an embryo, and it could shrink a full 50% in the preserving fluids. Mall took it upon

=Development of the Auricle in the Human Embryo= [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]]

...d in a majority of gilts in which all of the uterus was removed except one embryo and its corresponding portion of uterine horn on the 12th day of pregnancy. The Corpus Luteum of Pregnancy As It Is in Swine. Carnegie Inst. Contr. Embryol. 5: 69.

...e supply. The third group wliich numbered among its protagonists Retzius ('93), rle Vos ('94), and Mandl ('95) held that the follicle is not in any way p ...etic ganglia. We have verified this for the sow in a dissection of a large embryo prepared in formalin. However, the important work of Gaskell, Langley and D

=A Presomite Human Embryo (Shaw) with Primitive Streak and Chorda Canal with special reference to the ...appreciation of the gift we have associated the name of Dr Shaw with this embryo.

...pg|90px|left]] This historic 1931 paper describes an early human embryo, [[Carnegie stage 8]]. =A Young Human Embryo (Embryo Dobbin) with Head-Process and Prochordal Plate=

==Peters's Embryo - Yolk-sac== ...that Peters's specimen has no allantois. In describing another very young embryo he had recorded that "as compared with the embryonic shield, the allantois

[[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]] ...vide and multiply as the embryonic cells. The same structures which in the embryo grow and multiply by a process of karyo- kinesis, in the adult absolutely f

...mative cells, i.e., all the cells enter directly into the formation of the embryo's body. ...organforming areas which later enter into the formation of the body of the embryo; auxiliary or non-formative tissue has no part in its composition. All coel

...ical study With few exceptions, the younger the stage of development of an embryo of a particular species the lower is the animal group which it resembles bo ...hology does not appear to be concerned with the further development of the embryo, although the genetic structure of the male gametes is of fundamental impor

...ed follicle, but which have usually completely disappeared by the time the embryo begins to implant in the uterus In some species true luteal cells are added (esold H I 11937 Ih gona lotrophic liorm 5 93-103

...e, it was nevertheless apparent that a venous injection of the body of the embryo was often produced, and the impression was gained that a communication exis ...om the mouth, reaching it by way of the Eustachian tube. Using, in the pig embryo, the heart as the mechanism for injecting the ink, extravasation from the c

...ernal appearance and dimensions suggest that it is a [[Carnegie stage 19]] embryo ([[Week 7]], 48 - 51 days, 16 - 18 mm). {{Carnegie stage 19 links}}

...s were fertilized and were in stages from the one-celled to the six-celled embryo. Five of the twentyfour sows mentioned were obtained before a method of dis ...ictured in her recent contribution to the early vasculogenesis of the pig (Carnegie Institution of Washington, Contributions to Embryology, No. 18, 1917).

ribs in a 30-mm. CR embryo, for some cervical and upper and of the tympanic annulus of a 42—mm. CR embryo by Mall

embryo (Heuser and Streeter, 1941 ; Hertig embryo extract prepared from 19- to 20day-old guinea pig embryos (Blandau and

Carnegie Institution of Washington ...n of data was made by the statistical staff of the Department of Genetics, Carnegie Institution of Washington (Cold Spring Harbor, Long Island), and by Mr. Wil

...unity to collect this material; and to the Department of Embryology of the Carnegie Institution for technical assistance. ...lantois is intimately applied. Nutritive substances passing from mother to embryo must then traverse both the uterine epithelium and the columnar chorionic e

...arily spherical in form. Spherical cells are comparatively numerous in the embryo, and in the adult the resting white blood corpuscles, which float freely in A, From a 7 mm. embryo; B, from one of 26 mm.; ch, chroma tin knot; g. s, ground substance; 1, lin

embryo without the cooperation of the female, and whether the result is male Swammerdam (1637-80), Haller (1708-77), Bonnet (1720-93) and Spallanzani (1729-99) believed that the pre-existing parts of the new

...s thesis by Stewart describes development of the blood supply to the human embryo basal ganglia. =The Development of the Blood Supply to the Human Embryo Basal Ganglia=

...the early development of the neural folds and sensory anlagen of the human embryo. ...ez and Evans’ (’25) significant monograph on “The development of the human embryo during the period of somite formation, including embryos with 2 to 16 pairs

| These are links to other normal Carnegie Collection numbered embryos available on this educational site. {{Carnegie numbered embryo links}}

...ing a number 0f [[Carnegie Collection]] embryos. Note in addition to the [[Carnegie Stages]] (horizons) the term "stage" is used within this paper to describe Department of Embryology, Carnegie Institution of Washington, Baltimore; Division of Neurologicall Surgery, Un

...ntary bone and does not articulate with the sternum, and finally the human embryo, where the clavicle reaches its fullest development. ...to 10.5 mm. are described in the paper now under review. The 13-mm. human embryo corresponds essentially to the 18-mm. pig and 13-mm. cat. The pericardial c

site of a developing ovum. (From Carnegie Institution, No. C467.) coUiculi. (From Carnegie Institution, No. C713.)

namely, is there a separate germinal plasm set apart in the early embryo which used to refer to those germ cells which possibly segregate early in the embryo,

structural relationships between the developing embryo and the uterus. These comprise a succession of stages of placental metabolic demands of the developing embryo and fetus.

Carnegie Institution Of Washington, Department Of Embryology, The Johns Hopkins Univ in the embryo is controlled by a hormone,

...tion of the interaction among the developing and growing organs within the embryo. The study of the growth influences of one embryonic . organ on another is ...and then continues without interruption until a free living larva or young embryo is formed. This then proceeds to grow and change until the adult structure

...hese were consequently regarded as the fundamental differentiations of the embryo, determining its subsequent history. Next, differentiations among the cleav ...zation of the ovum is that of polarity, aheady described (6.gf., Figs. 42, 93). In many cases the polarity of the ovum can be traced back into oogonial s

...yolk. This substance contains the principal foodstuffs for the developing embryo. Studies on the yolk of the hen’s egg indicate that it contains water (50 ...ter chapters that the animal pole marks the anterior end of the developing embryo and the vegetal pole marks the posterior end. There is also reason to belie

...ls sie stehenden Wirbelthieren, erseheinen aber aueh bei dem menselilieben Embryo nieht vor Ablauf der ersten beiden Monate nach der Bmpfangniss. ...it they communicate with the exterior by a common opening. In the 4.75-mm. embryo (fig. 7) the naso-hypophyseal invagination has shifted toward the dorsal si

...dly number of which have appeared from the Department of Embryology of the Carnegie Institution of Washington. ...generosity of Dr. Carl G. Hartman from the Department of Embryology of the Carnegie Institution. These three macaque monkeys (Macaca mulatta) were killed on th

...p. 1391. Corner, G. W. 1915 Corpus luteum of pregnancy as it is in swine. Carnegie ...ctomjr in relation to the secondary sex characters of some domestic birds. Carnegie Inst. Washington, 243. Hegar, K. 1910 Studien zur Histogenese des Corpus lu

...empt was made to destroy just enough tissue along the dorsal aspect of the embryo to insure complete elimination of the neural-crest material and leave the v ...rophotographically in figures 1 and 2, which are taken from sections of an embryo of the chick (14) ^ which was subjected to operation at the close of the se

...presents the direction of the first outgrowth of the cochlear pouch of the embryo. As shown by Streeter ('07) for the human, this first growth of the cochlea and the acoustic and facial nerves in the human embryo. Am. Jour.

...ir in the female and the XY pair in the male. In this ani afighe members 0i 93°11 Pair are usually found together as indiTHE SIGNIFICANCE OF MEIOSIS 37 ...act that it was once thought that a very large portion of each side of the embryo always originated from this ring in a manner to be described below (see con

the embryo of the mouse and rabbit is lower embryo, until the sprouting of the primary

Washington, D. C. Published by the Carnegie Institution of Washington Carnegie Institution Of Washington, Publication No. 142

WASHINGTON, D. C. Published by the Carnegie Institution of Washington 1911 Carnegie Institution of Washington, Publication No. 142

...) stated that there are present in the developing islet cells of the sheep embryo minute safranophile granules. These have since been observed by Laguesse (' Fat droplets may also occur in islet cells. Dogiel ('93) noticed certain corpuscles which stained black in osmic-acid solution; he

...h he believed was the essential element in that it contained the preformed embryo in an intangible way. That is, the sperm animalcule of the ram contains a l ...rged capsule is soon formed which assumes the size and shape of the future embryo at the time of hatching (fig. 123). (See Tavolga, '50.) In the brook lampre

90-93 days the development of the embryo.

’93). A. In a salmon embryo after Furst. The position of

...ble percentage of fats. This important fact has been determined by Krehl ('93), Rosenfeld ('01), Leick ...n known in skeletal muscle for a long time. Knoll ('89-'91) and Schaffer ('93) described the dark fibers as containing manj^ interstitial granules and fa

From the Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland ...ation of the cavities in the cartilaginous capsule of the ear in the human embryo. Amer. Jour. Anat., vol. 22.

III. Description of a swell mechanism in the head of Testudinata. ... 93 A. The musculus constrictor venae jugularis internae 93

Translation by Joat V Nonidu Carnegie Institution Wuhington Translation by Jos6 P. Nonidez Carnegie Institution of Washington

University of Cincinnati Carnegie Institution {{Ref-Carey1921a}} Nine figures 93

Aided by the Carnegie Institution. ...us structures in the wall of the cerebral vesicle and neural tube of a cat embryo undoubtedly relate to mitochondria.

...o him the problem: If the spermatic fluid might stimulate the heart of the embryo in the process of fertilization, why might not other fluids produce the sam ...the diverse modifications which it undergoes, all the other organs of the embryo. '

...that described for other hemopoietic organs, e.g., yolk-sac of 10-mm. pig embryo, 5 yolk-sac of mongoose embryos, 6 and red bone-marrow. 7 ...of mesenchymal 'angioblasts' in the living blastoderm of the two-day chick embryo grown in Locke's solution, by which the blood-vessel lumen forms. But these

...ATION OF THE CAVITIES IN THE CARTILAGINOUS CAPSULE OF THE EAR IN THE HUMAN EMBRYO== Department of Embryology, Carnegie Institution of Washington, Baltimore,