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[[File:Stage_13_image_077.jpg|thumb|300px|Early embryonic gall bladder ([[Carnegie stage 13]], [[Week 4]])]] ...clear. In this study, we performed a DiI fate-mapping analysis using whole-embryo cultures of mouse early somite-stage embryos. Here, we show that the majori

...hilly 1987|link=Embryology History - Ronan O'Rahilly|Ronan O'Rahilly (1987 Carnegie Labs)]] ...t study,<ref name=Weller1933>{{Ref-Weller1933}}</ref> used the following [[Carnegie Collection]] embryos: stage {{CS9}} (No. {{CE1878}}), {{CS10}} ({{CE391}};

...is historic 1914 paper by Waterston describes a very early embedding human embryo. ...negie stage 12]] (26 - 30 days), caudal neuropore closes, Somite Number 21-29.

...dividual serial slices have also been incorporated into a 3D model of this embryo. * Stage 13 - Week 4, 26 - 30 days, 3 - 5 mm, Somite Number 21 - 29

This is the final Carnegie stage of embryonic development in Week 8. After this development is conside See also [[Carnegie stage 23#Events|'''Carnegie stage 23 Events''']]

PLoS One. 2010 Jul 29;5(7):e11877. * Stage 3 - embryo presence of somites and optic vesicles (epiboly was one-half complete) 96 d

...sion]] and [[Movie_-_Model_Embryo_to_128_Cell_Stage|Flash version]]. Added Carnegie collection [[Carnegie_stage_8#Carnegie_Collection|stage 8 images]]. ...- Model Embryo to 32 Cell Stage‎‎|Quicktime version]] and [[Movie - Model Embryo to 32 Cell Stage‎‎|Flash version]].

...]]) 17:55, 22 December 2014 (EST) Added [[Model Embryo 7.5mm Movie 1|Human Embryo 7.5mm model]] (Stage 15) from the [[Blechschmidt Collection]]. Added draft ...ll]] ([[User talk:Z8600021|talk]]) 19:38, 15 December 2014 (EST) Added new embryo images [[Carnegie_stage_13#Hill_Collection|Stage 13]] and [[Carnegie_stage_

--[[User:S8600021|Mark Hill]] 00:26, 29 October 2010 (UTC) [[Prostate Development]] and [[Urinary Bladder Developme --[[User:S8600021|Mark Hill]] 05:29, 9 October 2010 (UTC) Updated [[In Vitro Fertilization]] with information a

...New York (Dec 27-29, 2016). Describes the vitelline arteries in the human embryo. Modern Notes: [[Carnegie stage 10]] | [[week 4]]

The "Mateer Embryo" was later catalogued as Carnegie Embryo {{CE1399}} Carnegie stage {{CS8)). ...Poor || Formol || P || Trans. || 10 || {{HE}} etc. || 1916 || "Mateer embryo" described by Streeter (1920)

# Embryo staging and collections File:BrauneB1.jpg|Plate 29 and 30. Uterus and Fetus Position at Term

1. What period of human development (in weeks) do the 23 Carnegie stages cover? 3. At what Carnegie stage does the human neural tube normally completely close?

...) cushions (left hand panel). The right hand panel, which is from the same embryo but sectioned in short axis, shows how the left lateral atrioventricular cu '''Fig. 28.''' The images are from a mouse embryo at [[:Category:Mouse E15.5|E15.5]]. Septation is now complete. The short ax

...ke this a [[Carnegie stage 10|stage 10]] or [[Carnegie stage 11|stage 11]] embryo. =Photographs of a Series of Sections of an Early Human Embryo=

[[File:Stage 22 image 209.jpg|thumb|300px|Human embryo ([[Carnegie stage 22|stage 22]]) [[Sensory_-_Smell_Development|nasal epithelium]] devel File:Frazer1910 fig01.jpg|embryo 5 mm

...l|Maternal changes video]] and table of [[Fetal_Development#Carnegie_Fetal|Carnegie embryos at fetal stage]]. [[User:Z8600021|Mark Hill]] ([[User talk:Z8600021|talk]]) 10:00, 29 July 2017 (AEST) Added 2016 Science [[2016Lecture-Gametogenesis-Fertilizat

--[[User:Z8600021|Mark Hill]] 17:24, 29 December 2012 (EST) [[Zygote]] page updated with pronuclei information. ...[[Assisted_Reproductive_Technology#Assisted_Reproductive_Technology_.28USA.29|USA ART 2009 Report graphs]] also added to [[USA Statistics]].

...2, 1989). There is also a newer 1993 Downs and Davies staging of the mouse embryo included in each Theiler stage (Staging of gastrulating mouse embryos by mo * One-cell stage embryo (fertilised egg)

...lopment. Start by looking briefly at the overview of the Carnegie stage 13 embryo GIT from one end to the other. Then work through the listed specific serial sections of the embryo identifying the GIT features. Alternatively step through the serial section

...lar layer separating ectoderm and endoderm, mesoderm also lies outside the embryo as '''extra-embryonic mesoderm''' (covered in placenta lecture). Embryonic | {{Embryo logocitation}}

...ula; fusion of chorio-amniotic folds, chorio-amniotic stalk; neural plate; embryo bent dorsally; bud of allantoic stalk ...y; ectochorionic cyst collapsing; allantoic stalk projects into exocoelom; embryo bent dorsally

...the [[Harvard Collection|Harvard Embryological Collection (H.E.C.)]] and [[Carnegie Collection]]. ...were prepared from the following embryological series: 2.4, 4, 6.3, 8, 12, 29, and 40 mm.

...gnancy embryos. This was the very first publication in the series from the Carnegie Institution of Washington called [[Book_-_Contributions_to_Embryology|Contr =On The Fate Of The Human Embryo In Tubal Pregnancy=

1.What period of human development (in weeks) do the 23 Carnegie stages cover? 3.At what Carnegie stage does the human neural tube normally completely close?

[[File:Stage_13_image_077.jpg|thumb|300px|Early embryonic gallbladder ([[Carnegie stage 13]], [[Week 4]])]] ...the hepatocardiac channel. During expansion into the transverse septum at Carnegie Stage (CS)12 the liver bud develops as two dorsolateral lobes or 'wings' an

'''1. What period of human development (in weeks) do the 23 Carnegie stages cover?''' '''3. At what Carnegie stage does the human neural tube normally completely close?'''

[[Gastrointestinal_3D_stage_13_Movie|Week 4-5 embryo (stage 13) Movie]] | valign="top" | '''Begin by observing the internal structure of the embryo at the end of week 4 and the beginning of week 5.'''

[[File:Human Carnegie stage 10-23.jpg|thumb|300px|Carnegie Embryos]] ...collection numbering also incorporated the Blechschmidt embryo collection (Carnegie Nos. 10315-10434 ) in 1972, the collection embryos have now been returned t

...ark_Hill.jpg|50px|left]] This historic 1938 paper describes the "Falkiner" embryo, named after one of the co-authors. See also about this embryo: {{Ref-Falkiner1932}}

...etimes referred to as the Minot Collection, now forms part of the larger [[Carnegie Collection]]. The collection was described in detail by Minot (1905).<ref n [[Carnegie Collection]] - HDAC 7 Charles Sedgwick Minot Embryological Collection

[[File:Stage11_sem4.jpg|thumb|300px|Human head ([[Week 4]], [[Carnegie stage 11|Stage 11]]) showing buccopharyngeal membrane breakdown.]] ...during the process of gastrulation from the {{endoderm}} of the trilaminar embryo (week 3) and extends from the {{buccopharyngeal membrane}} ({{oral membrane

[[File:Kyoto940 stage21-07.jpg|thumb|150px|Kyoto embryo (940) [[Carnegie stage 21|stage 21]] histology]] ...:Z8600021|talk]]) 16:03, 1 November 2016 (AEDT) Added Kyoto embryo (940) [[Carnegie stage 21|stage 21]] histology images.

...}</ref> Later in 1921 along with Mall published a review of abnormal human embryo development.<ref>{{Ref-Mall1921}}</ref> ...lips of the blastopore (in the late gastrula stage) to other parts of the embryo and found that as expected they differentiated into structures characterist

| [[File:Human embryo day 2.jpg|200px|Stage 2 Day 2]] | [[File:Human embryo day 3.jpg|200px|Stage 2 Day 3]]

The embryo is now 1.0 - 1.5 mm in size. {{Carnegie stage 8 links}}

...h, liver, and dorsal pancreas primordia differentiated during {{CS12}} (21-29 somites) and CS13 (≥ 30 somites). The differentiation of four pairs of ph Human embryo small intestine secondary loops (week 7 to 8).{{#pmid:26297675|PMID26297675

| Blastocyst (ICM apparent) 16-40 compacted cells. Zona pellucida present. Embryo progresses from morula to the blastocyst. Early evidence of the blastocoeli ...ial arch become prominent. The preotic sulcus is visible in the 2-3 somite embryo. The cardiogenic plate begins to form and the foregut pocket is clearly vis

...ric 1955 paper by Mckay and co-authors describes human [[Carnegie stage 14|Carnegie horizon (stage) 14]] embryos. Currently only a brief abstract is included o '''Modern Pages:''' [[Carnegie stage 14]] | [[Week 5]] | [[Embryology History - Arthur Hertig|Arthur Herti

Embryo Liverpool II was later described in - {{Ref-HarrisonJeffcoate1953}} =A Presomite Human Embryo showing an early stage of the Primitive Streak=

...the [[Harvard Collection|Harvard Embryological Collection (H.E.C.)]] and [[Carnegie Collection]]. ...rrence in man were established by the investigations of Wilson and Anson (’29, two publications; twelve cases), and in mammals generally by those of Hoff

...to age fetuses based upon their bone ossification using embryos from the [[Carnegie Collection]]. ‘This is the average C-R length of all fetuses whose stated age was 29 or 3 months and indicates the basis for setting up the grouping in GR age.

...ng the ""Biggart" [[Carnegie stage 7]] and the younger "Macafee" embryo [[Carnegie stage 5]]. {{Carnegie stage 5 links}}

...blood development, including the fact that the red corpuscles in the early embryo are nucleated and that the later cells lack a nucleus. Specific informatio ...red blood cells begins to be evident during the second month in the human embryo and (2) that few nucleated red cells are found by the middle of the third m

...and, hence, the earliest axial structure presently known in the mammalian embryo may thus be functionally involved in shaping extraembryonic membranes and, ===Carnegie Stages Comparison Table===

[[File:Canine embryo E35-38 image003.jpg|thumb|Canine Embryo (E35-38)]] ...cularly high. So far, no puppy has been obtained from an in vitro-produced embryo. In contrast, cloning of somatic cells has been used successfully over the

...ine editor has replaced the traditional Roman numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

[[File:Stage11_sem4.jpg|thumb|300px|Human head ([[Week 4]], [[Carnegie stage 11|Stage 11]]) showing buccopharyngeal membrane breakdown.]] ...process of gastrulation from the [[E#endoderm|endoderm]] of the trilaminar embryo (week 3) and extends from the [[B#buccopharyngeal membrane|buccopharyngeal

...icle in the splendid collection of serial sections of human embryos in the Carnegie Laboratory of Embryology at the Johns Hopkins Medical School} My purpose wa ...to acknowledge the courtesy extended me by Dr. George L. Streeter, of the Carnegie Laboratory of Embryology, in placing the facilities

! width=120px|Carnegie Stage * 27 to 29 weeks - 17 % had not descended to the scrotum

[[File:Stage11_sem4.jpg|thumb|300px|Human head ([[Week 4]], [[Carnegie stage 11|Stage 11]]) showing buccopharyngeal membrane breakdown.]] ...during the process of gastrulation from the {{endoderm}} of the trilaminar embryo (week 3) and extends from the {{buccopharyngeal membrane}} to the {{cloacal

...indicate the absence of pouches. Weller (30) described a two somite human embryo which according to his description possessed the first pharyngeal pouch. Th Corner (4) described the foregut of a 10-somite human embryo, as being compressed dorso-ventrally with the anterior end immediately unde

...les of the development of the systemic Jymphatic vessels in the manmialian embryo. Anat. Rec, vol. 4, no. 2, pp. 399-423. ...n the domestic cat. Am. Journ. Anat., vol. 6, Abstr. Anat. Rec, vol. 1, p. 29.

{{Carnegie No.20 Header}} through the lateral canal of a rabbit embryo (fig. 457, page 735), in which this

...of the human embryo between Carnegie stage 19 to 23 in week 8 using the [[Carnegie Collection]] embryos. {{Carnegie stage 19 links}}

...ine editor has replaced the traditional Roman numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

...Fig. 17 . 168; F ig. 18. 29 ; Fi l(. 19. B 12~; !'ig. 20u. 1:3; Fi". 20b. 29.\ ; Fig. 20c. (;2 : Fig. 20d . 9S. ...alous stapes is the result: the l1ol'sesl1(ie-sllapetl stapes in a l-l-mm. embryo is an example of this defect (fig. 7).

...uld be clearly timed in the mouse and found in the literature on the human embryo. ...lation and fertilization times were unascertainable so that the age of the embryo is determined by the mating time plus or minus 30 minutes. The time for mat

{{Streeter-embryo to fetus}} ===Week 26 to Week 29===

==Appendix 1 - Embryos In The Carnegie Collection== The Carnegie specimens of stages 2-23 are listed in the following tables.

...portion of the pancreas. The study include embryos and fetuses from the [[Carnegie Collection]]. =The Development of the Islands of Langerhans in the Human Embryo=

...his historic 1945 paper by Shaner describes a Carnegie Stage {{CS9}} human embryo of two to three pairs of somites. {{Carnegie stage 9 links}}

[[Carnegie stage 12]] * Characteristic feature: 21–29 pairs of somites

...d the development of the thoracic vertebrae using human embryos from the [[Carnegie Collection]]. [[Embryology History - Charles Bardeen|Charles Bardeen]] | [[Carnegie Embryos]]

...crown-rump length of 14.5 mm. The embryo was in Streeter’s Horizon early [[Carnegie stage 18|XVIII]]. Several factors led to the selection of this particular h ...arious levels in a plane coinciding with the transverse plane in which the embryo was sectioned. Figure 3 illustrates the internal structure that is evident

[[File:Stage_22_image_217.jpg|thumb|300px|Cerebrum development human embryo (week 8, Stage 22)]] | {{Embryo logocitation}}

...e 19|stage 19]]; {{CE1584}} [[Carnegie stage 19|stage 19]]; {{CE1535}} [[Carnegie stage 23|stage 23]]. '''Modern Notes:''' {{embryonic}} | {{CRL}} | [[Carnegie Collection]]

'''University of Vienna''' - Institute of Histology and Embryology Embryo Collection ...27 pairs of somites [[Carnegie stage 12]] in [[Week 4]] (Somite pairs 21 - 29)

...er birth. This development generates the most complex structure within the embryo and the long time period of development means in utero insult during pregna ...ormation of the caudal neural tube to the tip of the caudal portion of the embryo was finished at [[Carnegie_stage_17|stage 17]]. The postcloacal gut had com

...c 1957 paper by O'Rahilly is a description of the development of the human embryo limb cartilage. ...4; Hagen, 1900; Lewis, 1902; Griifenberg, I905; Hesser, 1926). In a 27—mm. embryo, Schulin (1879) found that all the skeletal elements of the hand were ehond

...0 series and 12 dissected tonsillar regions from the [[Carnegie Collection|Carnegie Institution, Department of Embryology]], and 50 series and 19 dissected ton ...tance. I also wish to acknowledge the generous help of Dr. G. L. Streeter, Carnegie Institution of Embryology, in placing at my disposal abundant material.

...by Odgers describes an early embryo development, later characterised as [[Carnegie stage 8]]. {{Carnegie stage 8 links}}

...Mall describes the human embryos in the collection that would become the [[Carnegie Collection]]. There is also a [[:File:1904 - Catalogue of the collection of [[Carnegie Collection]] | [[Carnegie Embryos]]

...ould make clear the significance of the unfamiliar topography of the young embryo and give us a perspective from which to follow out the internal changes occ ...dult anatomical form it is, nevertheless, the beginning of the body of the embryo. The remaining regions of the blastocyst will take part only in the formati

...s to hold me. Before I was born out of my mother generations guided me. My embryo has never been torpid, nothing could overlay it.''" - Walt Whitman, Song of ...The tub in which we imagine the tree planted simulates the pool which the embryo has excavated for itself in the wall of the mother's uterus (Fig. 14, p. 58

{{Carnegie No.20 Header}} ...ilaginous capsule of the ear undergoes during its development in the human embryo are accomplished in part by a progressive and in part by a retrogressive di

...ix of [[Carnegie stage 18|stage 18]], [[Carnegie stage 19|stage 19]] and [[Carnegie stage 21|stage 21]] embryos. {{Carnegie stage 18 links}}

both at the Carnegie Institute and at the Rockefeller Institute. He is an honorary member and fe Carnegie Institution of Washington Department of Embryology, Baltimore

...ove can be seen on the head surface before fusing to enclose the duct in [[Carnegie stage 19]]. J ouves (’97) studied the sheep and man, and found in a 19 mm. human embryo both lacrimal ducts present but without any connection with the surface epi

...ht, Sitting Height, Head Size, Foot Length, and Menstrual Age of the Human Embryo= Published by the Carnegie Institution of Washington Washington, 1920

...2 paper by Patten describes early development of heart in the {{chicken}} embryo. ...another as far as could be determined, were available for the WOI'l{. One embryo in each of these sets was reserved for study as a cleared and stained entir

...his paper appear to be from the [[Carnegie Collection]] though some of the embryo number, CRL and ages do not appear to match the catalogue information. Fig. l. — Embryo 87; size, 25 mm crown-rump length; age, about 8 weeks. The capsule is mostl

...istoric histological study of the development of the meninges of the human embryo spinal cord. Our current understanding of interstitial cell development and ...by the Department of Embryology, sincere appreciation is expressed to the Carnegie Institution of Washington, and to Dr. G. W. Corner, Director. The author is

...pment of the trachea and esophagus and includes several embryos from the [[Carnegie Collection]]. Department Of Embryology, Carnegie Institution Of Washington, Baltimore, Maryland

...are not Carnegie stages, use the embryo CRL to approximately convert to [[Carnegie Stages]]. ...bryology_15|Historic - Urogenital Development]] | [[Carnegie Embryos]] | [[Carnegie Collection]]

...describes the development of the human sympathetic nervous system using [[Carnegie Collection]] embryos: {{CE460}}. ...cated, were made on human embryos included in the [[Carnegie Collection|'''Carnegie Embryological Collection''']]. It is a real pleasure to express my indebted

...Mark_Hill.jpg|90px|left]] This 1935 paper by Gilbert describes early human embryo hypophysis (pituitary) development. ...hms, ’32; Gilbert, ’34). These investigations have shown that in the early embryo the ventral surface ectoderm of the head is closely adherent to the floor

[[Carnegie stage 14]] ...ation in the size of the chorion for a given stage than in the size of the embryo. That the growth of the chorion is much influenced by environmental nourish

...ube with two dilatations: one represents a ruptured chorionic sac with its embryo still inside: the other sac was unruptured, entirely distinct from the firs ...a tubal pregnancy described by Mall ('15) and ]\Ieyer ('20), listed in the Carnegie collection as no. 825 (fig. 2). Externally the tube bore a single swelling

...blood, excrete waste, reabsorb water and have endocrine functions. In the embryo, there are several stages in their development closely linked to genital de | {{Embryo logocitation}}

|[[File:Mark_Hill.jpg|50px|left]] This historic 1956 paper describes using [[Carnegie Collection]] early human development in week 2 and 3. [[Carnegie Embryos|Carnegie Embryos in this paper]]: {{CE8698}} | {{CE8794}} | {{CE8663|}} | {{CE8663}

...omach, liver, and dorsal pancreas primordia differentiated during CS12 (21-29 somites) and CS13 (≥ 30 somites). The differentiation of four pairs of ph Human embryo small intestine secondary loops (week 7 to 8).{{#pmid:26297675|PMID26297675

...rnegie stage 7]] (26 - 30 days), caudal neuropore closes, Somite Number 21-29. {{Carnegie stage 7 links}}

...ody. The head and neck are one of the most complicated structures that the embryo forms, with special intermediate structures (the {{pharyngeal arch}}) and c ...cavity floor02.jpg|thumb|alt=Human embryo Stage 13 oral cavity floor|Human embryo pharyngeal arches (Stage 13 oral cavity floor)]]

...bers the embryos examined in sequence 1-30, but also gives Mall's Carnegie embryo numbers in the tabulation. Carnegie Embryo {{CE186}}

.... In 1959, Khvatov found a pronuclear egg in a serially sectioned oviduct (29). In 1965, Dickmann, Clewe, Bonney, and Noyes studied, by means of phase mi ...entations in those portions immediately adjacent to the polar bodies (fig. 29). The plasma membrane was smooth and devoid of typical microvilli. A few bl

...r and heart grow rapidly forming obvious external swellings on the ventral embryo surface. The liver's initial embryonic function is mainly {{cardiovascular} ...man_Embryology_17-7|1912 Liver]] | [[Paper - Functions of the liver in the embryo|1920 Embryonic Function]] | [[Book_-_Text-Book_of_Embryology_12#The_Develop

...ioned and stained. Of the embryos and fetuses studied, 152 belonged to the Carnegie Institution Department of Embryology. The remaining specimens were from the ...s to Dr. G. L. Streeter for the use of the Embryo1ogical Collection of the Carnegie Institution and supplying figures 15, 16, and 17. I also wish to thank Pro

[[Carnegie stage 15]] ...in [[1987 Developmental Stages In Human Embryos - Stage 14|stage 14]], the embryo resulting from these influences is bilaterally flattened, having a curved o

Human embryo of 23 somites occurs at [[Carnegie stage 12]] in [[Week 4]]. ==Description of a Human Embryo of Twenty-three Paired Somites==