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...embryo catalogue.jpg|thumb|alt=Orts Llorca Madrid embryo catalogue|Madrid embryo catalogue]] The human embryo histology collection was started in 1935 by Professor Orts-Llorca (1905-199

See also [[Carnegie stage 22#Events|'''Carnegie stage 22 Events''']] {{Carnegie stage 22 links}}

...1996. There are also sets of [[Carnegie stage 22 - serial sections]] and [[Carnegie stage 22 - selected serial sections]]. ...ks:''' [[Carnegie stage 13]] | [[Carnegie stage 22 - serial sections]] | [[Carnegie stage 22 - selected serial sections]]

! | [[Carnegie stage table|'''Carnegie Stages''']] | [[File:Stage20_bf1c.jpg|200px|link=Carnegie stage 20]]

Mouse embryo, Theiler stage 21. * 52-55 somite pairs

...hn Rock|John Rock]] and included embryos that were incorporated into the [[Carnegie Collection]]. [[Carnegie Collection]] Embryo No. {{CE8004}} (stage 5b) from 1945 paper.<ref name=Hertig1945a>{{Ref-Hert

...n either age or size. The human embryonic period proper is divided into 23 Carnegie stages. Criteria beyond morphological features include age in days, number ...an take from as little as 10 days in chickens to nearly 60 days in humans. Carnegie is the name of a historical US Institute that historically categorised thes

The historic "Dobbin" embryo was named after Dr Roy Dobbin (Cairo, Egypt) who originally provided the sp This [[Carnegie stage 8]] human embryo was 960 µm long and was subsequently serially sectioned and published in a

[[Media:BGD2010-Embryo Lab 170510-605.mp3|listen Part 5]] | [[:File:BGD2010-Embryo Lab 170510-605.mp3|download]] (1.8 Mb MP3 13:14) [[File:Stage19-_limb_rotation.jpg|thumb|Human Embryo (Stage19) limb rotation]]

| Carnegie No. 763 || 8? | Carnegie No. 1399 || 7

...re are also sets of [[Carnegie stage 22 - selected serial sections]] and [[Carnegie stage 13 - serial sections]]. | [[:File:Stage 22 image 055.jpg|55]]

[[File:Kyoto940 stage21-07.jpg|thumb|150px|Kyoto embryo (940) [[Carnegie stage 21|stage 21]] histology]] [[User:Z8600021|Mark Hill]] ([[User talk:Z8600021|talk]]) 12:55, 6 November 2016 (AEDT) Updated [[Prenatal Diagnosis]] with information abo

...ctures/ilectures.lasso?ut=153&id=27393 Lecture Audio - 25 Sep 2008 - 13:00 55 min] === Stage 13/14 Embryo ===

This is the final Carnegie stage of embryonic development in Week 8. After this development is conside See also [[Carnegie stage 23#Events|'''Carnegie stage 23 Events''']]

--[[User:Z3218792|Gabriela Pinget]] 13:55, 13 August 2009 (EST) ''What period of human development (in weeks) do the 23 Carnegie stages cover?''

...int Louis University, Saint Louis, Missouri, and Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland ...identified by a number of external and internal characteristics, and each embryo at a given stage has a similar degree of organization and differentiation t

[[File:Carnegie_Institute_of_Washington_logo.jpg|thumb|Carnegie Institute of Washington]] ...The papers documented not only early human development, using mainly the [[Carnegie Collection]] of embryos, but also that in animal models of development.

...System Development]] added [[:File:Human- Stage 22 integument 01.jpg|human embryo stage 22 skin image]] and movie of melanoblast migration in muse skin [[Qui --[[User:S8600021|Mark Hill]] 01:55, 21 September 2010 (UTC) [[Integumentary System - Tooth Development]] stage

...]]) 17:55, 22 December 2014 (EST) Added [[Model Embryo 7.5mm Movie 1|Human Embryo 7.5mm model]] (Stage 15) from the [[Blechschmidt Collection]]. Added draft ...ll]] ([[User talk:Z8600021|talk]]) 19:38, 15 December 2014 (EST) Added new embryo images [[Carnegie_stage_13#Hill_Collection|Stage 13]] and [[Carnegie_stage_

| Blastocyst (ICM apparent) 16-40 compacted cells. Zona pellucida present. Embryo progresses from morula to the blastocyst. Early evidence of the blastocoeli ...ial arch become prominent. The preotic sulcus is visible in the 2-3 somite embryo. The cardiogenic plate begins to form and the foregut pocket is clearly vis

...24, inclusive, and‘ 22 and 19, are taken from drawings made of the Keibel embryo, which is at present not available for study. figure 27, the lowest of the In the Mall embryo the orifices of the two ureters and the Wolffian duct are, so far as I was

...y Models]] page with information about [[Embryology_Models#Carnegie_Models|Carnegie Models]]. ...o_Embryology_Carnegie_Institution_No.56|Contributions to Embryology Vol.12 Carnegie Institution No.56 (1921)]].

| The bilaminar ({{epiblast}} and {{hypoblast}}) embryo is now about 0.2 mm diameter in size. The three extra embryonic spaces (amn {{Carnegie stage 6 links}}

...sion]] and [[Movie_-_Model_Embryo_to_128_Cell_Stage|Flash version]]. Added Carnegie collection [[Carnegie_stage_8#Carnegie_Collection|stage 8 images]]. ...- Model Embryo to 32 Cell Stage‎‎|Quicktime version]] and [[Movie - Model Embryo to 32 Cell Stage‎‎|Flash version]].

[[File:Carnegie stages species comparison.jpg|500px]] ...the sheep uterus and infect the conceptus trophectoderm in a transspecies embryo transfer model'''{{#pmid:20610723|PMID20610723}}

The embryo is now 1.0 - 1.5 mm in size. {{Carnegie stage 8 links}}

...2, 1989). There is also a newer 1993 Downs and Davies staging of the mouse embryo included in each Theiler stage (Staging of gastrulating mouse embryos by mo * One-cell stage embryo (fertilised egg)

Embryo Liverpool I was previously described in - {{Ref-HarrisonJeffcoate1953}} {{Carnegie stage 7 links}}

{{Carnegie Collection stage 23 table}} ...[:Category:Carnegie Embryo 7425|No 7425]]. Lower row, [[:Category:Carnegie Embryo 4570|No. 4570]]. All views are at the same magnification.

...and, hence, the earliest axial structure presently known in the mammalian embryo may thus be functionally involved in shaping extraembryonic membranes and, ===Carnegie Stages Comparison Table===

...ula; fusion of chorio-amniotic folds, chorio-amniotic stalk; neural plate; embryo bent dorsally; bud of allantoic stalk ...y; ectochorionic cyst collapsing; allantoic stalk projects into exocoelom; embryo bent dorsally

...etimes referred to as the Minot Collection, now forms part of the larger [[Carnegie Collection]]. The collection was described in detail by Minot (1905).<ref n [[Carnegie Collection]] - HDAC 7 Charles Sedgwick Minot Embryological Collection

...ine editor has replaced the traditional Roman numeral used in the text for Carnegie staging with a number link to that stage online information. {{Carnegie stage table 1}}

[[File:Stage11_sem4.jpg|thumb|300px|Human head ([[Week 4]], [[Carnegie stage 11|Stage 11]]) showing buccopharyngeal membrane breakdown.]] ...process of gastrulation from the [[E#endoderm|endoderm]] of the trilaminar embryo (week 3) and extends from the [[B#buccopharyngeal membrane|buccopharyngeal

...the [[Harvard Collection|Harvard Embryological Collection (H.E.C.)]] and [[Carnegie Collection]]. ...were studied; of those between 3.0 and 6.6 mm., forty-four series in the [[Carnegie Collection]]. The series in the Harvard Collection which best displayed the

...c 1957 paper by O'Rahilly is a description of the development of the human embryo limb cartilage. ...4; Hagen, 1900; Lewis, 1902; Griifenberg, I905; Hesser, 1926). In a 27—mm. embryo, Schulin (1879) found that all the skeletal elements of the hand were ehond

[[File:Human Carnegie stage 10-23.jpg|thumb|300px|Carnegie Embryos]] ...collection numbering also incorporated the Blechschmidt embryo collection (Carnegie Nos. 10315-10434 ) in 1972, the collection embryos have now been returned t

...human embryo|Human Embryo vascular development (week 8, stage 20 Carnegie Embryo No. {{CE460}})]] ...embryo 50 mm. long (Carnegie Collection, No. 96)|Human embryo 50 mm long (Carnegie Collection, No. {{CE96}}.]]

...}</ref> Later in 1921 along with Mall published a review of abnormal human embryo development.<ref>{{Ref-Mall1921}}</ref> ...lips of the blastopore (in the late gastrula stage) to other parts of the embryo and found that as expected they differentiated into structures characterist

...0 series and 12 dissected tonsillar regions from the [[Carnegie Collection|Carnegie Institution, Department of Embryology]], and 50 series and 19 dissected ton ...tance. I also wish to acknowledge the generous help of Dr. G. L. Streeter, Carnegie Institution of Embryology, in placing at my disposal abundant material.

[[Carnegie stage 8]] ...: the primitive pit, the notochordal canal, and the neurenteric canal. The embryo is presomitic, i.e., somites are not yet visible.

...the ossicles in the middle ear were independent in different locations. At Carnegie Stage 17 a homogeneous interzone clearly defined the incus and malleus anla Cross-section of human embryo [[Carnegie stage 22]] during [[Week 8]].

...icle in the splendid collection of serial sections of human embryos in the Carnegie Laboratory of Embryology at the Johns Hopkins Medical School} My purpose wa ...to acknowledge the courtesy extended me by Dr. George L. Streeter, of the Carnegie Laboratory of Embryology, in placing the facilities

| {{Embryo citation}} * Teenagers the rate of NTDs decreased by almost 55%

{{Carnegie No.20 Header}} ...ilaginous capsule of the ear undergoes during its development in the human embryo are accomplished in part by a progressive and in part by a retrogressive di

both at the Carnegie Institute and at the Rockefeller Institute. He is an honorary member and fe Carnegie Institution of Washington Department of Embryology, Baltimore

==Appendix 1 - Embryos In The Carnegie Collection== The Carnegie specimens of stages 2-23 are listed in the following tables.

[[Carnegie stage 6]] ...ncidence of sex chromatin in the trophoblast and chorionic mesoblast of an embryo (No. {{CE7801}}) of stage 6, and in the chorionic mesoblast and umbilical v

...are not Carnegie stages, use the embryo CRL to approximately convert to [[Carnegie Stages]]. ...bryology_15|Historic - Urogenital Development]] | [[Carnegie Embryos]] | [[Carnegie Collection]]

...genesis of the thyroid follicles (Norris, ’16), has been carried on at the Carnegie Institute of Embryology and at the University of Minnesota under the superv This study is based upon the collection of human embryos in the Carnegie Institute of Embryology at Baltimore and upon those in the Anatomical Labor

...and Sotelo and Porter (40), and in the rabbit by Zamboni and Mastroianni (55). The fine morphology of the human penetrated ovum has never been investiga ...tion they clump, and a nuclear membrane becomes reconstituted around them (55). At the end of this process, a female pronucleus is formed. Mean- while, t

...evelopment of the limbs, body-wall and back using human embryos from the [[Carnegie Collection]]: {{CE2}}, {{CE12}}, {{CE22}}, {{CE43}}, {{CE76}}, {{CE106}} {{ ...Figs. F to I, Plates VI to IX, are based upon wax-plate reconstructions of Embryo XXII.

...to age fetuses based upon their bone ossification using embryos from the [[Carnegie Collection]]. ...11 67 6 30 21 12 3 71 7 48 24 21 16 109 8 10 I 1 11 5 37 9 5 2 1 4 12 10 ‘ 55 33 20 16 124 Total 194 116 80 63 Grand total 310 143 -453Terata ’ 5 7 3 2

...omandibular joint, that connecting the mandible to the skull. Pater uses [[Carnegie Collection]] embryos: {{CE1318}}, {{CE1455}}, {{CE3990}}, {{CE5652}}, {{CE6 ...ular nt in 5 human fetuses, the smallest one having a crown—rump length of 55 mm. He observed in 3 of these specimens that some of the fiber bundles of t

...Mall describes the human embryos in the collection that would become the [[Carnegie Collection]]. There is also a [[:File:1904 - Catalogue of the collection of [[Carnegie Collection]] | [[Carnegie Embryos]]

...Mark_Hill.jpg|90px|left]] This 1935 paper by Gilbert describes early human embryo hypophysis (pituitary) development. ...hms, ’32; Gilbert, ’34). These investigations have shown that in the early embryo the ventral surface ectoderm of the head is closely adherent to the floor

...and others on human vertebral column development using embryos from the [[Carnegie Collection]].{{#pmid:517765|PMID517765}}{{#pmid:7216919|PMID7216919}}{{#pmi ...-resolution digitized imaging data (n = 34) obtained in human embryos with Carnegie stage (CS) between {{CS17}} and {{CS23}}. The rib cage became detectable as

...The origin of the mesenteric lymph sac in the pig. Anat. Rec. vol. 2, pp. 55-57. ...les of the development of the systemic Jymphatic vessels in the manmialian embryo. Anat. Rec, vol. 4, no. 2, pp. 399-423.

[[File:Low1909 plate01fig01.jpg|thumb|Plate 1 fig1 Model human embryo 18 mm mandible viewed from above]] ...ale as separate elements. More recently H. Fuchs states that in the rabbit embryo there is a distinct and separate splenial element, that the condylar cartil

...evelopment of the limbs, body-wall and back using human embryos from the [[Carnegie Collection]]: {{CE2}}, {{CE12}}, {{CE22}}, {{CE43}}, {{CE76}}, {{CE106}} {{ ...Figs. F to I, Plates VI to IX, are based upon wax-plate reconstructions of Embryo XXII.

...infiltrated decidua. That these chorionic vesicles both once contained an embryo, no embryologist will doubt. It is evident also that both were retained for ...of the thorax. The small nodule, which represents the only remnant of the embryo belonging to the larger chorionic vesicle, contains a large cavity, with th

|[[File:Mark_Hill.jpg|50px|left]] This historic 1956 paper describes using [[Carnegie Collection]] early human development in week 2 and 3. [[Carnegie Embryos|Carnegie Embryos in this paper]]: {{CE8698}} | {{CE8794}} | {{CE8663|}} | {{CE8663}

...e free to use our judgment in methods of fixation and preservation. If the embryo is perfectly fresh or possibly living, we use, of course, the most refined ...straight and other measurements and weights also are taken. The age of the embryo is estimated on the basis of weight, crown-rump, and foot length, and the e

...|841]], [[:Category:Carnegie Embryo 5537|5537]], and [[:Category:Carnegie Embryo 6521|6521]]). Note Mall's reference is cited as 1913, but it is the 1912 he ...eveloping septa in accord with Tandler’s findings. Moreover, the age of the embryo fits nicely into the Tandler series and also with the stages described in

...|90px|left]] This historic 1929 paper by Ingalls (1880-1949) describes a [[Carnegie Collection]] human embryos segmental thickenings in the dorsal ectoderm of ...(726), appear as shown in figure A. This represents a left lateral view of embryo no. 155, C.R. 11.8 mm. The thickenings or dises are indicated by the row of

...ht, Sitting Height, Head Size, Foot Length, and Menstrual Age of the Human Embryo= Published by the Carnegie Institution of Washington Washington, 1920

* '''Contributions to Embryology''' - [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington Series]] A historic series of papers published by the Carnegie Institution of Washington early in the 20th Century.

...ogy|Carnegie Institution of Washington - Contributions to Embryology]] | [[Carnegie Collection]] The Embryological Collection of the Carnegie Institution of Washington presents splendid opportunities for the investiga

Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland, and Department of Zoology, ...rvated by the oculomotor. Bonnet (1901) likewise found, in a 16-somite dog embryo, a pair of mesodermal condensations derived from a medial mass of cells at

Department of Embryology, Carnegie Institution of Washington, Baltimore, Mn. ...th its associated tiny mass of protoplasm, which alone will make the chick embryo, becomes all but lost on the surface of the yolk. But that speck of protopl

...e 19|stage 19]]; {{CE1584}} [[Carnegie stage 19|stage 19]]; {{CE1535}} [[Carnegie stage 23|stage 23]]. '''Modern Notes:''' {{embryonic}} | {{CRL}} | [[Carnegie Collection]]

=Notes on Irregularities of Ovogenesis and Abnormal Development of the Embryo in Cavia= ...f deviations from the normal in the early stages in the development of the embryo of Cavia. These specimens were obtained during a series of investigations o

By A. M. Hain (Carnegie Research Fellow), Institute of Animal Genetics, University of Edinburgh, ...were largely defrayed by grants from the Medical Research Council and the Carnegie Trust for the Universities of Scotland.

...d the development of the thoracic vertebrae using human embryos from the [[Carnegie Collection]]. [[Embryology History - Charles Bardeen|Charles Bardeen]] | [[Carnegie Embryos]]

....jpg|90px|left]] This historic 1941 paper by Gilmour describes early human embryo blood formation. ....065 x 0.045 mm. Age about 16 days, probably slightly younger than Peters’ embryo (1899).

Development of this system commences in the embryo, continues through the fetal period then with key changes around birth, onl ...Reproductive development has a long maturation timecourse, begining in the embryo and finishing in puberty. (More? [[Puberty Development]])

The measurements of the embryo are as follows: C.R., crown-rump or sitting height; C.H., crown—hee1 or s <div id="Carnegie Embryo 6"></div>

...{CE460}} is classified as a [[Carnegie stage 20]] (week 8, {{GA}} week 10) embryo. The following pages also relate to this topic. =The Cartilaginous Skull of a Human Embryo Twenty-One Millimeters in Length=

...ube with two dilatations: one represents a ruptured chorionic sac with its embryo still inside: the other sac was unruptured, entirely distinct from the firs ...a tubal pregnancy described by Mall ('15) and ]\Ieyer ('20), listed in the Carnegie collection as no. 825 (fig. 2). Externally the tube bore a single swelling

...bellum of Man. He stated that the rhombic lip begins to form, in the human embryo, at the beginning of the fifth week, and that it extends from the cervical ...gave an illustration of a transverse section through the hind-brain of an embryo rat (3 mm. long) which certainly closely resembled some of the figures give

...ix of [[Carnegie stage 18|stage 18]], [[Carnegie stage 19|stage 19]] and [[Carnegie stage 21|stage 21]] embryos. {{Carnegie stage 18 links}}

...pg|90px|left]] This historic 1931 paper describes an early human embryo, [[Carnegie stage 8]]. =A Young Human Embryo (Embryo Dobbin) with Head-Process and Prochordal Plate=

...y Atlas of the 13-mm. Pig Embryo. (Prefaced by younger stages of the chick embryo.) The Wistar Institute Press, Philadelphia, iv & 104 pp. ...tatus of the germ-cell problem in vertebrates. Biol. Rev., Vol. 20, pp. 45-55.

...of the lower axial skeleton and lower limbs using human embryos from the [[Carnegie Collection]]: {{CE2}}, {{CE22}}, {{CE45}}, {{CE62}}, {{CE109}}, {{CE144}}, [[Embryology History - Charles Bardeen|Charles Bardeen]] | [[Carnegie Embryos]]

...indicate the absence of pouches. Weller (30) described a two somite human embryo which according to his description possessed the first pharyngeal pouch. Th Corner (4) described the foregut of a 10-somite human embryo, as being compressed dorso-ventrally with the anterior end immediately unde

...a constant invitation to theorizing. Since it is well known that the human embryo does not normally pass through any developmental stage comparable to either ...stages in the normal development of the cloacal region of the young human embryo and shows that the primordia of the genital tuberele are at first paired. O

{{Carnegie stage 7 links}} =The Chorion and Endometrium of the Embryo H.R.1=

=Chapter III. The Human Embryo= ==Calculation of the Age of the Human Embryo==

...[[Book_-_Contributions_to_Embryology_Carnegie_Institution_No.55|1920 Human Embryo Head Size]] | [[Book_-_Contributions_to_Embryology_Carnegie_Institution_No. File:Low1909 fig01.jpg|embryo 18 mm

...icken}} pulmonary vessel development. [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]] ...this kind some obstacles are sure to be encountered, even in so simple an embryo as the chick. In mammalian embryos these are harder to overcome and offer a

...to tissues that are lost during development. It includes several of the [[Carnegie Collection]] embryos in the figures. ...thology, Harvard Medical School, Boston, and the Department of Embryology, Carnegie Institution of Washington, Baltimore.

=Development of the Auricle in the Human Embryo= [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]]

...al origin, are very early differentiated from the otic vesicle in the 7 mm embryo as a medial diverticular projection (Bast, Anson and Gardner, ’47). This ...and Gardner (’47) pointed out that the sac overlies the sinus in the 50 mm embryo.

'''Human Embryo - developing cortex''' * [[Week 8]], [[Carnegie stage 22]].

...l the cytoplasm is divided into two giving nse to a two cell st ige of the embryo or ovum (Fig 28B and C) Each of the daughter cells contains an equal number ...stage stage F —eight cell stage ( \fter Lev is and Hartman 1933 ) of tf e Carnegie Institution of Washington y c *00

'''Human Embryo - developing cortex''' * [[Week 8]], [[Carnegie stage 22]].

...and mouse models.<ref name="PMID2297894"><pubmed>2297894</pubmed></ref> As embryo grows, these receptors mature, axons ascend and are restricted to the derma |Merkel cells are derivatives of the epidermis of the developing embryo. They are able to be seen, with short dendrites, as early as week 8 in embr

...ovary after differentiation. The germinal ridge is first established in the embryo 9 days p.c., and the gonads are definitely difierentiated into ovaries or ...ed with the aid of a planimeter, and the average of the two gonads of each embryo was plotted on the accompanying graph (text-fig. 1). It can be seen from th

...- Blastocyst (ICM apparent) 16-40 compacted cells. Zona pellucida present. Embryo progresses from morula to the blastocyst. Early evidence of the blastocoeli ...blast induces distal visceral endoderm (DVE) formation that will establish embryo anterior–posterior axis.{{#pmid:15302604|PMID15302604}}

...90px|left]] [[Historic Embryology Papers]] | [[Embryonic Development]] | [[Carnegie Collection]] =Cyclopia in the Human Embryo=