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...lar layer separating ectoderm and endoderm, mesoderm also lies outside the embryo as '''extra-embryonic mesoderm''' (covered in placenta lecture). Embryonic | {{Embryo logocitation}}

Department of EmbryologyThe Carnegie Institution Of Washington, Baltimore, Maryland ...empt was made to recover a specimen comparable to the youngest known human embryo.

...a description of early {{twinning}}. The embryo is classified as Carnegie Embryo No. {{CE1522}}. Department of Embryology, Carnegie Institution of Washington

...ola. Human Embryology and Teratology. (1990) Lippincott Williams & Wilkins 400 pages ...The future dorsal foramina (containing dorsal rami) face laterally in the embryo and are in line with the thoracicolumbar intervertebral foramina. Considera

[[File:Stage11_sem6.jpg|thumb|300px|Human embryo (Week 4, 24 days, Carnegie stage 11, 13 somite pairs)]] ...ic tissues and systems are beginning to differentiate) from the trilaminar embryo. With many parallel processes, descriptions begin to get complicated! Many

| width=420px|<html5media height="600" width="400">File:Stage20 MRI S02.mp4</html5media> [[File:Kyoto Embryo Collection - cover.jpg|200px|link=http://itunes.apple.com/us/book/id1143922

[[File:Carnegie stages species comparison.jpg|500px]] ...mid:20615540|PMID20615540}} "Sheep preantral follicles (PFs) measuring 250-400 microm in diameter were cultured for six days in serum-free media supplemen

! 1910-30 Carnegie Stages | [[File:Human Carnegie stage 10-23.jpg|600px|link=Carnegie Stages]]

! 1910-30 Carnegie Stages | [[File:Human Carnegie stage 10-23.jpg|600px|link=Carnegie Stages]]

[[Carnegie stage 1]] ...a. 0.1 mm) and weight (ca. 0.004 mg) of the organism at fertilization, the embryo is "''schon ein individual-spezifischer Mensch''" (Blechschmidt, 1972). The

...ht, Sitting Height, Head Size, Foot Length, and Menstrual Age of the Human Embryo= Published by the Carnegie Institution of Washington Washington, 1920

Department of Embryology, Carnegie Institution of Washington ..., with all parts in their natural connection and the contours of the Whole embryo and of the different organs clearly recognizable. On the other hand, the la

...cess of implantation and early differentiation of cells that will form the embryo and the placenta. ...to all material derived from this fertilized zygote and includes both the embryo and the non-embryonic tissues (placenta, fetal membranes).

...in the embryo as at birth, but the deformities of the head and neck of the embryo are of such a nature that it can not survive long enough to admit of compar ...xomphaly. Other anomalies, however, are more difficult to recognize in the embryo as sharply defined malformations.

* Embryo 1 - 17th day of the menstrual cycle, ovulation probably occurred less than * Embryo 2 - 16th day of the cycle; coitus had taken place 2 and 4 days previously.

[[File:Stage_22_image_217.jpg|thumb|300px|Cerebrum development human embryo (week 8, Stage 22)]] | {{Embryo logocitation}}

[[File:Human Carnegie stage 10-23.jpg|thumb|Carnegie Embryos]] [[File:Streeter02.jpg|thumb|Measuring embryo size]]

...in the embryo as at birth, but the deformities of the head and neck of the embryo are of such a nature that it cannot live long enough to admit of comparison ...r, are more difficult to recognize as sharply defined malformations in the embryo.

...parturition) is approximately 64 days and there have been identified about 400 congenital disorders relating to dog development. Many of these development [[File:Canine embryo E35-38 image003.jpg|thumb|Canine Embryo (E35-38)]]

...by Odgers describes an early embryo development, later characterised as [[Carnegie stage 8]]. {{Carnegie stage 8 links}}

...r ear. These human embryos are [[Carnegie Embryos]] and fetuses from the [[Carnegie Collection]]. ...f the Scala Tympani, Scala Vestibuli and Perioticular Cistern in the Human Embryo=

...}</ref> Later in 1921 along with Mall published a review of abnormal human embryo development.<ref>{{Ref-Mall1921}}</ref> ...lips of the blastopore (in the late gastrula stage) to other parts of the embryo and found that as expected they differentiated into structures characterist

=The Development of the Venous Sinuses of the Dura Mater in the Human Embryo= Department of Embryology, Carnegie Institution of Washington

...rain vascular development includes descriptions of many embryos from the [[Carnegie Collection]] including: {{CE84}}, {{CE96}}, {{CE144}}, {{CE199}}, {{CE234a} ...Developmental Alterations in the Vascular System of the Brain of the Human Embryo=

[[Carnegie stage 5]] ...asures approximately 0.1-0.2 mm in diameter. The significant dimensions of Carnegie specimens of stage 5 are listed in Table 5-1. The external and internal dia

...10) the right side of the neck and thorax was cut in sagittal sections. In embryo (9) and the six foetuses the neck and upper part of the thorax were cut in (10) Embryo, 9th-10th week, no measurement recorded, sagittal sections at 15 9, of righ

...e are no other data known to me. LiUie expresses the following opinion (p. 400): "A number of mammalian groups could be at once excluded from consideratio ...ube with two dilatations: one represents a ruptured chorionic sac with its embryo still inside: the other sac was unruptured, entirely distinct from the firs

...genesis of the thyroid follicles (Norris, ’16), has been carried on at the Carnegie Institute of Embryology and at the University of Minnesota under the superv This study is based upon the collection of human embryos in the Carnegie Institute of Embryology at Baltimore and upon those in the Anatomical Labor

{{Carnegie stage 7 links}} =The Chorion and Endometrium of the Embryo H.R.1=

...ix of [[Carnegie stage 18|stage 18]], [[Carnegie stage 19|stage 19]] and [[Carnegie stage 21|stage 21]] embryos. {{Carnegie stage 18 links}}

The measurements of the embryo are as follows: C.R., crown-rump or sitting height; C.H., crown—hee1 or s <div id="Carnegie Embryo 6"></div>

...id variation in the human embryo|The supracondyloid variation in the human embryo]]. (1934) Anat. Rec. 314-329. =The Supracondyloid Variation in the Human Embryo=

...statistics of Williamson would indicate that it may be found but once in 2,400 cases." Williams adds, however, that in his own experience it occurred even ...belonging very largely in the later months of pregnancy, while that in the Carnegie Collection, on the other hand, belongs very largely in the earlier months.

...paper by Goss describes early development of heart contraction in the rat embryo. ...of beginning contraction, a Wax plate reconstruction Was prepared of a rat embryo of corresponding age fixed in the uterus.

...at first slowly, then more rapidly, until it was finally taken over by the Carnegie Institution of Washington in 1915. ...ter the collection had been transferred to the Carnegie Institution, about 400 specimens were collected in one year. It will be observed also that approxi

...opment. Here, a detailed step-by-step protocol for extended ex utero mouse embryo culture is provided. The ability to grow normal mouse embryos ex utero from ...nner-cell contacts in the ICM, which activates Oct4 in the preimplantation embryo. Oct4 is highly expressed but unstable at E3.25-LNC, and stabilizes at high

...e, in which it opened in the naso-pharynx. In the development of the human embryo we see these three stages reproduced.<ref> See Professor J. E. Frazer, Lanc ...(Dev. of Palate) ; E. Fawcett, Journ. Anat. and Physiol. 1906, vol. 40, p. 400 (Ossific. of Palate). See also references, p. 159,</ref> At the end of the

...This 1916 paper by Cunningham in the [[Book_-_Contributions_to_Embryology|Carnegie Institution of Washington - Contributions to Embryology]] series describes =On the Development of the Lymphatics of the Lungs in the Embryo Pig=

| {{Embryo logocitation}} # Development of the '''indifferent gonad''' - (genital ridge) early embryo

...e show that although blood cell formation in its early stages in the human embryo and especially the yolk sac follows the same general lines as in other mamm ...y with the magma strands of the chorionic cavity. At the caudal end of the embryo these yolk sac mesodermal cells, which will give rise to the yolk sac mesen

...s human embryo collection that will eventually form the beginning of the [[Carnegie Collection]]. ...Developmental Alterations in the Vascular System of the Brain of the Human Embryo]]. (1921) [[Book_-_Contributions_to_Embryology#Volume_VIII|Contrib. to Embr

[[File:Rabbits.png|right|400 px]] '''Brief timeline of rabbit embryo model use'''

| [[File:Logo.png|alt=Embryo logo|80px]] | {{Embryo citation}}

...pment of the trachea and esophagus and includes several embryos from the [[Carnegie Collection]]. Department Of Embryology, Carnegie Institution Of Washington, Baltimore, Maryland

...University of Chicago collection that were eventually contributed to the [[Carnegie Collection]]. ...would approximately between [[Carnegie stage 11]] (13 - 20 somites) and [[Carnegie stage 12]] (21 - 29 somites).

...he caudal end of the spinal cord in human embryos using embryos from the [[Carnegie Collection]]. {{Carnegie Collection fetal table}}

...foetus. Edinburgh. , 1904. Manual of antenatal pathology and hygiene. The embryo. Edinburgh. DANDY, W. E., 1910. A human embryo with seven pairs of somites measuring about 2 mm. in length. Amer. Jour. An

...e free to use our judgment in methods of fixation and preservation. If the embryo is perfectly fresh or possibly living, we use, of course, the most refined ...straight and other measurements and weights also are taken. The age of the embryo is estimated on the basis of weight, crown-rump, and foot length, and the e

...e pulli in ovo (London: Joannem Martyn, 1673).|Cover Illustration: Chicken embryo, from M. Malp Dissertatio epistolica de formatione pulli in ovo (London: Jo Cover Illustration: Chicken embryo, from M. Malp Dissertatio epistolica de formatione pulli in ovo (London: Jo

=A Human Embryo Before the Appearance of the Myotomes= ...ed investigation of its more essential features, especially as regards the embryo proper, has been undertaken. The extra-embryonic structures, chorion, body-

By A. M. Hain (Carnegie Research Fellow), The Institute of Animal Genetics, Edinburgh University, ...y defrayed by grants (to A.M.H.) from the Medical Research Council and the Carnegie Trust for the Universities of Scotland.

...lar pregnancy, gestational trophoblastic disease) using embryos from the [[Carnegie Collection]]. Some of these concepts are historic and have been updated wit ...nks:''' [[Abnormal_Development_-_Hydatidiform_Mole|Hydatidiform_Mole]] | [[Carnegie Collection]]

...90px|left]] [[Historic Embryology Papers]] | [[Embryonic Development]] | [[Carnegie Collection]] =Cyclopia in the Human Embryo=

...onare Geschlechtzellen’ in the intestinal epithelium of a four~weeks human embryo. The supposed sex cells were disposed in such a way as to suggest an active ...considered to be germ cells in the lateral plates of mesoderm of a 2.3—mm. embryo, and as these plates were folded under the gut in 2.8-mm. embryos, the germ

...y Atlas of the 13-mm. Pig Embryo. (Prefaced by younger stages of the chick embryo.) The Wistar Institute Press, Philadelphia, iv & 104 pp. Corner, G. W., 1915. The corpus luteum of pregnancy as it is in swine. Carnegie Inst., Contrib. to E-mbryoL, Vol. 2, pp. 69-94.

...use of the intrinsic nature of the subject, for the functions of which the embryo or fetus is capable at various times are determined by the growth of the ne ...representation, then, allows us to observe the general growth picture from embryo to adult, and gives us a basis upon which to establish a more detailed anal

...ch populate the cytoplasm adjacent to the pronuclei. g, Golgi complex. X 5,400. ...emely important because it implies that the mitochondrial endowment of the embryo is exclusively maternal in derivation.

...lation for 20 days did not maintain corpora lutea in intact lambs (up to 2,400 I.U. prolactin per day) or in lambs hypophysectomized on the day of ovulati ...d in a majority of gilts in which all of the uterus was removed except one embryo and its corresponding portion of uterine horn on the 12th day of pregnancy.

Acting Lecturer and Demonstrator in Anatomy, University College, London; tate Carnegie Research Fellow in Embryology, and Assistant in the Depariment of Anatomy, This stage is represented by one embryo 25 mm. in length, the cranial extremity of which has been reconstructed in

=A Human Embryo of Twenty-Four Pairs of Somites= ...] | [[Book_-_Contributions_to_Embryology|Contributions to Embryology]] | [[Carnegie stage 12]] | [[Week 4]]

...ants from the Penrose Fund of the American Philosophical Society, from the Carnegie Corporation of New York and from the University of Pittsburgh. ...ndogenous stimulation and remarked that if he had been able to observe the embryo under more favorable conditions before its circulation had been disturbed,

Koelliker (’7 9) has noted in a rabbit embryo of eleven days a close relation between the anterior end of the notochord a caudal surface of Rathke’s pouch. In a 4.5-mm. embryo he saw

Carnegie Institution of Washington ...n of data was made by the statistical staff of the Department of Genetics, Carnegie Institution of Washington (Cold Spring Harbor, Long Island), and by Mr. Wil

...an study. This embryo (Teacher-Bryce Ovum No. 1) was later classified as [[Carnegie stage 6]] {{Carnegie stage 6 links}}

embryo without the cooperation of the female, and whether the result is male enlarged compartment where the egg or developing embryo may be retained.

structural relationships between the developing embryo and the uterus. These comprise a succession of stages of placental metabolic demands of the developing embryo and fetus.

Department of Embryology, Carnegie Institution of Washington, Baltimore, Maryland ...ovulationem, leaving only five and one-half days‘ actual development of the embryo to birth. The rate of development is compared with Eutherian mammals.

...some way this morphology of the egg is . related to the morphology of the embryo developed from the egg, and hence is called its promorphology. ,, ...materials in general related with the vegetative organs of the developing embryo.

...cribes the structure of the adult brain-stem. See also the series from the Carnegie Institution of Washington called [[Book_-_Contributions_to_Embryology|Contr Washington, D. C. Published by the Carnegie Institution of Washington 1914

...dly number of which have appeared from the Department of Embryology of the Carnegie Institution of Washington. ...elucidation of cytological features at a magnification greater than 300 or 400 times or by the application of special fixatives and differential stains.

| These are links to other normal Carnegie Collection numbered embryos available on this educational site. {{Carnegie numbered embryo links}}

Washington, D. C. Published by the Carnegie Institution of Washington Carnegie Institution Of Washington, Publication No. 142

WASHINGTON, D. C. Published by the Carnegie Institution of Washington 1911 Carnegie Institution of Washington, Publication No. 142

Aided by the Carnegie Institution. ...us structures in the wall of the cerebral vesicle and neural tube of a cat embryo undoubtedly relate to mitochondria.

...yolk. This substance contains the principal foodstuffs for the developing embryo. Studies on the yolk of the hen’s egg indicate that it contains water (50 ...ter chapters that the animal pole marks the anterior end of the developing embryo and the vegetal pole marks the posterior end. There is also reason to belie

...o him the problem: If the spermatic fluid might stimulate the heart of the embryo in the process of fertilization, why might not other fluids produce the sam ...the diverse modifications which it undergoes, all the other organs of the embryo. '

...that described for other hemopoietic organs, e.g., yolk-sac of 10-mm. pig embryo, 5 yolk-sac of mongoose embryos, 6 and red bone-marrow. 7 ...of mesenchymal 'angioblasts' in the living blastoderm of the two-day chick embryo grown in Locke's solution, by which the blood-vessel lumen forms. But these

...) stated that there are present in the developing islet cells of the sheep embryo minute safranophile granules. These have since been observed by Laguesse (' Pankreas beim nienschlichen Embryo. Arch, mikros. Anat., Bd. 64. Kyrle, J. 1908 Ueber die Regenerationsvorgang

...f the Scala tympani, Scala vestibuli and perioticular cistern in the human embryo. Nine figures 299 Embryo 12.84

A. In a salmon embryo after Furst. The position of the cell body. They share in its trophic functions, as is nerve of an embryo of

....40 per cent at 200 grams, 0.37 per cent at 300 grams and 0.35 per cent at 400 grams. ...59 per cent at 200 grams, 0.55 per cent at 300 grams, and 0.54 per cent at 400 grams.

a. Chromosomes in the embryo 3 a. Chromosomes in the embryo

...ATION OF THE CAVITIES IN THE CARTILAGINOUS CAPSULE OF THE EAR IN THE HUMAN EMBRYO== Department of Embryology, Carnegie Institution of Washington, Baltimore,

...rth day after operation from infection in the wound, having lost meanwhile 400 g. in weight. ...IC SAC AND ITS TOPOGRAPHICAL RELATION TO THE TRANSVERSE SINUS IN THE HUMAN EMBRYO

...bryos were arranged accord- ing to measurement rather than age. The oldest embryo of the first series was 9.4 mm. in length, and the later series were select ...al cavity and in the pharynx, buds that resemble mature buds of the oldest embryo studied in all essential details except size. The later maturing of the tas

...presents the direction of the first outgrowth of the cochlear pouch of the embryo. As shown by Streeter ('07) for the human, this first growth of the cochlea and the acoustic and facial nerves in the human embryo. Am. Jour.

...ennent have made experiments in which the paternal influence in the hybrid embryo was diminished. Tennent states that in the cross between Hipponoe and Toxop ' Tennent, Publication 132, Carnegie Institution, 1910.

Translation by Jos6 F. Nonidez Carnegie Institution of Washington ...reas, intervening sarcosomes. Flemming fixation, iron-hematoxylin stain. X 400. (The photomicrographs for figs. 11, 12, and 13 were made by Mr. William S.

Embryo: nine to fourteen days’ incubation 12 ...is il ne se distingue plus du restant de 1 ’epithelium p4ritoneal (’20, p. 400).

...ogy, With An Appendix On The Arteries And Veins In A Thirty Millimeter Pig Embryo== Ordinarily, the student is shown two dimensions of a piece of tissue or embryo, and left to imagine the third. Though whole mounts of chick embryos are ha

are comparatively numerous in the embryo, and in the adult the resting A, From a 7 mm. embryo; B, from one of 26 mm.; ch,

Staff Member, Department of Embryology, Carnegie Institution Of Washington. Baltimore. Maryland 400

Mabel Bishop. The nervous system of a two-headed pig embryo. Twenty figures 379 400

...esearch rooms" at Woods Hole wishes to acknowledge his indebtedness to the Carnegie Institution for the opportunities thus presented for the carrying on of thi 0.400

The youngest embryo in which any of the air-sacs appear as In the same embryo may be seen the first indication of the

...tion of the marsupial blastocyst with the trophoblast of the Eutheria, the embryo of the former, therefore, being without trophoblastic covering. ...on in similar conditions independently discovered by Patterson CIO) in the embryo of the Tatusia.

0.400 Carnegie Institution of Washington

...erning certain cytological characteristics of the erythroblasts in the pig embryo and the origin of non-nucleated erythrocytes by a process of cytoplasmic co ==The Development Of The Rectum In The Human Embryo==

...e take place, as is shown in the caudal end of the Wolffian body of a deer embryo of 6.4 mm. (fig. 3), where the increased growth of one limb of the transver ...uricular canal, and the adult relations reproduce essentially those of the embryo. Hence their comihon histological characters in the adult and the intimate

Howard Brown Stough. Modified mitosis in the chick embryo. Eight plates (sixty figures) 535 1. The embryo of the chick develops in its own egg entirely apart from the mother. For th