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- 20:49, 23 October 2017 Z5076158 talk contribs uploaded File:Isthmic Organiser.png (The role of the Isthmic Organiser. "The establishment of gene expression domains along the anterior-posterior axis helps to segment the developing brain into the forebrain, midbrain (MB), hindbrain (HB) and spinal cord (SC). The Isthmic Organizer (IsO;...)