From Embryology
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Embryology - 25 May 2020    Facebook link Pinterest link Twitter link  Expand to Translate  
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A personal message from Dr Mark Hill (May 2020)  
Mark Hill.jpg
I have decided to take early retirement in September 2020. During the many years online I have received wonderful feedback from many readers, researchers and students interested in human embryology. I especially thank my research collaborators and contributors to the site. The good news is Embryology will remain online and I will continue my association with UNSW Australia. I look forward to updating and including the many exciting new discoveries in Embryology!


This contains links to content pages that have been recently updated or added to this site. Since software updated to MediaWiki 1.26.2 and the total number of page views magicword no longer supported. Therefore cannot continue to provide view count updates from 2016 onwards. Individual pages will also no longer show "number of views" in the footer. Please note that not all site updates are logged on this page.

UNSW Embryology has now been available online and continuously updated for 20 years!

Current total number of topic pages - 6,243

10 Popular Pages  
Mark Hill.jpg

OK these are not my rankings, but by you the viewer.

These are the top ten popular pages based upon page views.

I have excluded the Main page, student projects and undergraduate course pages. Note, this data has not been updated since 1 January 2016, when page visit logs were removed.

1 January 2016 (EST)
  1. Embryonic Development‏‎ (399,216 views)
  2. Placenta Development‏‎ (183,296 views)
  3. Timeline human development‏‎ (154,106 views)
  4. Neural System Development‏‎ (116,916 views)
  5. Mouse Development‏‎ (115,361 views)
  6. Respiratory System Development‏‎ (112,977 views)
  7. Placenta - Abnormalities‏‎ (111,887 views)
  8. Implantation‏‎ (111,620 views)
  9. Fetal Development‏‎ (97,150 views)
  10. Chicken Development‏‎ (89,201 views)
Site Map - Main Page - expand to see pages 
Main Page Contributors Glossary Terms Categories Site map Site updates Recent changes New images
Editing Basics Images Tables Referencing Journal Searches Copyright Font Colours Virtual Slide Permalink My Preferences One Page Wiki Card Printing Movies Language Translation Student Movies Mobile Access Internet Browsers Moodle Navigation Short URLs
Medicine Foundations BGDA BGDB BGD2 Tutorial SH HM AE Med Projects
Science ANAT2341 ANAT2241 ANAT2511 ANAT3411 Sci Projects Editing Basics Textbooks Embryology Education
Development Overview Week 1 Week 2 Week 3 Week 4 Neural Mesoderm Gastrointestinal Tract Cardiovascular Vascular Head and Face Renal Genital Endocrine Embryo Stages Fetal Ultrasound Neonatal Postnatal Animal Movies
Medicine Audio Science Audio Gastrointestinal Tract Endocrine Heart Sounds
One Minute  
Human Development Human Timeline Human Embryo Endoderm Development Human Face and Palate
Carnegie Stages  
stage 1 stage 2 stage 3 stage 4 stage 5 stage 6 stage 7 stage 8 stage 9 stage 10 stage 11 stage 12 stage 13 stage 14 stage 15 stage 16 stage 17 stage 18 stage 19 stage 20 stage 21 stage 22 stage 23
Week 1 
(GA week 3)
stage 1 stage 2 stage 3 stage 4 Menstrual Cycle Fertilization Zygote Morula Blastocyst Meiosis Mitosis
Menstrual Cycle Oocyte Spermatozoa Twinning Genetic risk maternal age Trisomy 21 Trisomy 18 Trisomy 13 Hydatidiform Mole Lecture - Fertilization Lecture - Week 1 and 2
Week 2 
(GA week 4)
stage 5 stage 6 Implantation Trophoblast Human Chorionic Gonadotropin Pregnancy Test Twinning Lecture - Week 1 and 2
Week 3 
(GA week 5)
stage 7 stage 8 stage 9 Gastrulation Endoderm Mesoderm Ectoderm Placenta Coelomic Cavity Lecture - Week 3
Week 4 
(GA week 6)
stage 10 stage 11 stage 12 stage 13 Placodes Pharyngeal arches Lecture - Mesoderm Lecture - Ectoderm Lecture - Early Vascular
Week 5 
(GA week 7)
stage 14 stage 15 Placodes Pharyngeal arches Lecture - Head Atrial Ventricular Septation
Week 6 
(GA week 8)
stage 16 stage 17 Lecture - Head Outflow Tract
Week 7 
(GA week 9)
stage 18 stage 19 Lecture - Head Outflow Tract Heart Valves
Week 8 
(GA week 10)
stage 20 stage 21 stage 22 stage 23 Lecture - Head Outflow Tract Heart Valves
Cardiovascular Coelomic Cavity Endocrine Gastrointestinal Genital Head Immune Integumentary
Musculoskeletal Neural Neural Crest Placenta Renal Respiratory
Hearing Vision Smell Taste Touch
Genetic Environmental
Abnormal Systems  
Cardiovascular Coelomic Cavity Endocrine Gastrointestinal Tract Genital Head Integumentary Musculoskeletal Limb Neural Neural Crest Renal Respiratory Placenta Sensory Hearing Vision
Environmental Low Folic Acid Iodine Deficiency Nutrition Drugs Thalidomide Herbal Drugs Illegal Drugs Smoking Fetal Alcohol Syndrome TORCH
TORCH Infections Cytomegalovirus Hepatitis Virus HIV Parvovirus Polio Virus Rubella Virus Chickenpox Lymphocytic Choriomeningitis Virus Zika Virus Vaccination
Syphilis Gonorrhea Tuberculosis Listeria TORCH Infections
Zoonotic Toxoplasmosis Malaria Maternal Diabetes Maternal Hypertension Maternal Hyperthermia Maternal Inflammation Maternal Obesity Hypoxia Biological Toxins Chemicals Heavy Metals Radiation
Prenatal Diagnosis  
Pregnancy Test Amniocentesis Chorionic villus sampling Ultrasound Alpha-Fetoprotein Pregnancy-associated plasma protein-A Fetal Blood Sampling Magnetic Resonance Imaging Computed Tomography Non-Invasive Prenatal Testing Fetal Cells in Maternal Blood Preimplantation Genetic Screening Comparative Genomic Hybridization Genome Sequencing
Neonatal Diagnosis  
APGAR test Guthrie test Hearing test Electrocardiogram X-ray Tandem mass spectrometry ICD-10 DOHAD
XVII Congenital Malformations System Tables XVI Perinatal Period XV Pregnancy Childbirth
Axolotl Bat Cat Chicken Cow Dog Dolphin Echidna Fly Frog
Grasshopper Guinea Pig Hamster Kangaroo Koala Lizard Medaka Mouse Pig Platypus
Rabbit Rat Sea Squirt Sea Urchin Sheep Worm Zebrafish Life Cycles Timetable K12
Assisted Reproductive Technology
Stains Fixatives Menstrual Cycle Placenta Heart Liver Pancreas Gall Bladder Colon Renal Respiratory Histology Bone UNSW Histology
Histology Glossary: A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z | ANAT2241 Support | Histology | Histology Stains | Embryology Glossary
Virtual Slides
Historic Embryology Papers Historic Embryology Textbooks Embryologists
Historic Periods  
1700's 1800's 1900's 1910's 1920's 1930's 1940's 1950's 1960's 1970's 1980's Historic Papers
Historic Terminology
Human Embryo Collections  
Blechschmidt Collection Carnegie Collection Domenech-Mateu Collection Harvard Collection Hill Collection Hinrichsen Collection Hubrecht Collection Kyoto Collection Madrid Collection
Embryology Models DEC Information DEC Embryo Collections Embryologists
Carnegie - Contributions 17-18th C Anatomies Embryology Models
Historic Papers  
1800's 1900's 1910's 1920's 1930's 1940's 1950's Embryologists
Historic Periods  
1700's 1800's 1900's 1910's 1920's 1930's 1940's 1950's 1960's 1970's 1980's Historic Papers
iBooks Journals
Start Week 1 Week 2 Week 3 Week 4 Week 5 to 8 Fetus Arms and Legs
Heart Brain Growth Eyes and Ears Animals Humans and Animals Comparative Embryology Thalidomide Education
Genetics Epigenetics Mitosis Meiosis X Inactivation Signaling
hCG BMP Sonic hedgehog Homeobox FGF Nanog Notch Fox Pax
Retinoic acid Six Slit2/Robo1 Sox Tbx TGF-beta VEGF Wnt Hippo
Mouse Knockout microRNA Mechanisms Enhancers Protein Genetic Abnormal
World Population World Fertility World Infant Mortality Maternal Mortality Reports Australia Brazil Canada China Germany
India Indonesia Europe Myanmar Spain United Kingdom Romania Uganda United States
Australia’s mothers and babies  
2014 2013 2012 2011
2010 2009 2008 2007
Stem Cells  
Timeline Placental Cord Blood Adult | Induced Yamanaka Factors Somatic Cell Nuclear Transfer Ethics
Textbooks Education


Chicken Somitogenesis

February 2017

Mark Hill (talk) 14:08, 3 February 2017 (AEDT) Added Mouse Blastocyst Cdx2 Movie and Chicken Somitogenesis Movie.

List of abbreviations:
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25 May 2020

stage 14 SEM
Human embryo 18 mm mandible

January 2017

Mark Hill (talk) 11:35, 28 January 2017 (AEDT) Added collapsible site map (also above on this page).

Mark Hill (talk) 14:20, 27 January 2017 (AEDT) Added new virtual slide stage 14 SEM image.

Mark Hill (talk) 15:14, 9 January 2017 (AEDT) Added a series of historic papers by Ernest Frazer.

Mark Hill (talk) 13:02, 6 January 2017 (AEDT) Added historic papers including Low A. Further observations on the ossification of the human lower jaw. (1909) J Anat Physiol. 44(1): 83–95. PMID 17232830 Site includes many historic papers showing how our current understanding of embryonic development began.

Mark Hill (talk) 12:47, 2 January 2017 (AEDT) Updated Australian Statistics with NSW Mothers and Babies 2015 Report.

Mark Hill (talk) 05:06, 1 January 2017 (AEDT) Finalised 2016 statistics and updated footer and page citation for 2017.


Heart Movie (1951)
Chicken Movie (1961)

December 2016

Mark Hill (talk) 15:10, 13 December 2016 (AEDT) Added Heart Historic Movie 1951 animation with soundtrack describing heart septation and Historic Chicken Movie 1961.

Mark Hill (talk) 16:11, 1 December 2016 (AEDT) Added educational presentation UNSW Medicine Learning and Teaching Forum 2016 - Supporting Independent Learning with eBooks‎‎

Kyoto embryo (940) stage 21 histology

November 2016

Mark Hill (talk) 09:05, 26 November 2016 (AEDT) Australia’s mothers and babies 2014 summary added.

Mark Hill (talk) 15:06, 18 November 2016 (AEDT) Added Historic GIT papers Johnson FP. The development of the mucous membrane of the oesophagus, stomach and small intestine in the human embryo. (1910) Amer. J Anat., 10: 521-559. Johnson FP. The development of the rectum in the human embryo. (1914) Amer. J Anat. 16(1): 1-58.

Mark Hill (talk) 12:55, 6 November 2016 (AEDT) Updated Prenatal Diagnosis with information about cervical sampling of fetal trophoblast cells.

Mark Hill (talk) 16:03, 1 November 2016 (AEDT) Added Kyoto embryo (940) stage 21 histology images.

Hinrichsen ME16 (stage17) head

October 2016

Mark Hill (talk) 14:03, 23 October 2016 (AEDT) Added draft historic textbook. Arey LB. Developmental Anatomy. (1924) W.B. Saunders Company, Philadelphia.

Mark Hill (talk) 10:26, 19 October 2016 (AEDT) Added images

Hinrichsen Images: ME50 stage 12 | ME18 stage 13 | ME52 stage 14 | ME52 stage 14 Head | ME34 stage 16 | ME34 stage 16 Arches | ME16 stage 17 Head | ME16 stage 17 Nasal | S226 stage 18 | ME28 stage 19 | ME28 stage 19 Hand | ME54 stage 21 | ME54 stage 22 | ME45 stage 23 Head | ME45 stage 23 Face | Hinrichsen Collection

September 2016

Mark Hill (talk) 17:12, 12 September 2016 (AEST) Updated Renal System Development.

August 2016

Mark Hill (talk) 10:54, 31 August 2016 (AEST) Updated Molecular Development with molecular system links menu.

Mark Hill (talk) 12:50, 23 August 2016 (AEST) The Kyoto Collection eBook added as an external iBookstore link.

Mark Hill (talk) 12:05, 22 August 2016 (AEST) Added MRI movie Stage 13 pharyngeal arches and heart.

Mark Hill (talk) 14:38, 17 August 2016 (AEST) Added Stage 13 and Stage 14 movies

Mark Hill (talk) 11:05, 11 August 2016 (AEST) Updated Science ANAT2341 Lab 2.

Mark Hill (talk) 09:05, 10 August 2016 (AEST) Added Palate terms‎‎ and Hearing terms lists.

Mark Hill (talk) 00:34, 9 August 2016 (AEST) Added animal draft page Monito del Monte Development‎‎.

Mark Hill 21:32, 2 August 2016 Updated Hearing Development with embryonic stage information.

Mark Hill 16:12, 2 August 2016 Added new embryo stage animation.


Embryo stages 01.gif

MRI Stage 17 movie 1 icon.jpg

July 2016

Mark Hill 13:21, 22 July 2016 Updates Science ANAT2341 course 2016 and 2016 Course Outline PDF,

Mark Hill 12:23, 18 July 2016 Added Embryo stage 17 MRI movie.

Mark Hill 10:23, 16 July 2016 Help:Translation updated the language translation page.

2016 AIWH Folic Acid and Iodine Report

June 2016

Human Embryo Placental Blood Vessels

Mark Hill 11:03, 28 June 2016 Updated Folic Acid page with recent 2016 AIWH Report information.

Mark Hill 14:05, 23 June 2016 Added ultrasound movies abnormality detection for Velamentous Cord, Placenta Previa and Hypoplastic Left Heart Syndrome.

Mark Hill 20:55, 20 June 2016 Added draft historic papers on skull development Macklin CC. The skull of a human fetus of 40 mm 1. (1914) Amer. J Anat. 16(3): 317-386., Macklin CC. The skull of a human fetus of 40 mm 2. (1914) Amer. J Anat. 16(3): 387-426.

Mark Hill 02:43, 17 June 2016 Added draft historic textbook McMurrich JP. The Development Of The Human Body. (1914) P. Blakiston's Son & Co., Philadelphia, Pennsylvania. and new Main page news item.

Mark Hill 16:35, 14 June 2016 Added ultrasound movies abnormality detection for Dandy-Walker, Spina Bifida, Vasa Previa and Transposition Great Arteries.

Mark Hill 13:27, 13 June 2016 added historic paper drafts Bardeen Lewis 1901‎- Limbs, Whitehead and Waddell 1911 - sternum, Lisser 1911 - larynx, Pohlman 1911 - cloaca and Reagan 1912 - 5th aortic arch.

Mark Hill 10:23, 10 June 2016 Added stage 22 histology image gallery.

Mark Hill 22:55, 5 June 2016 Updated BGDA Practical - Placenta and Fetal Membranes.

organ of corti

May 2016

Mark Hill (talk) 19:01, 26 May 2016 (AEST) Added new Stage 18 Virtual Slides.

Mark Hill (talk) 12:25, 26 May 2016 (AEST) Added Heart terms.

Mark Hill (talk) 17:11, 24 May 2016 (AEST) Updated BGD2 Tutorial.

Mark Hill (talk) 14:09, 22 May 2016 (AEST) Updated Abnormal Development - Drugs‎.

Mark Hill (talk) 15:50, 20 May 2016 Updated Neural Crest Development‎‎.

Mark Hill (talk) 23:52, 18 May 2016 Hearing - Inner Ear Development‎‎ added images of organ of corti. Inner Ear Histology: image - cochlea | image - cochlear duct | image - organ of corti | image - organ of corn detaili | image - stria vascularis | Inner Ear | Histology

Mark Hill (talk) 20:55, 15 May 2016 Updated BGDB Lecture Face and Ear.

Mark Hill (talk) 10:13, 7 May 2016 (AEST) Updated Medicine BGDA Lecture 1 and BGDA Lecture 2.

Mark Hill (talk) 07:03, 5 May 2016 (AEST) Updated Medicine BGDA and BGDB practical templates.

17 to 23 somite human embryos
17 to 23 somite human embryos

April 2016

Statistics | Recent Changes | New Files | RSS Recent


Mark Hill (talk) 12:33, 27 April 2016 (AEST) Added One Minute Embryology a set of quick introductory videos with audio commentary designed to get students to begin to think about specific embryology topics. Also posted on UNSW theBox.

Mark Hill (talk) 10:06, 23 April 2016 (AEST) updated Monosomy page Cri du Chat syndrome information and Category:Terms.

Mark Hill (talk) 11:54, 22 April 2016 (AEST) Added historic reference templates.

Mark Hill (talk) 14:20, 21 April 2016 (AEST) Added historic paper Paper - The Anatomy of Human Embryos with Seventeen to Twenty-three Pairs of Somites‎

Mark Hill (talk) 08:03, 17 April 2016 (AEST) Updated Foundations Lecture - Introduction to Human Development and Foundations Practical - Introduction to Human Development for 2016.

Mark Hill (talk) 15:03, 5 April 2016 (AEST) Updated Wiki appearance to skin version 1.3.0 (2016-03-08), this has a clearer simplified layout with the navigation menu now located at the top of the page.

Spleen structure cartoon
Spleen structure cartoon

March 2016

Statistics | Recent Changes | New Files | RSS Recent

--Mark Hill (talk) 14:59, 29 March 2016 (AEDT) Updated K12 Comparative Embryology‎‎ for teacher development class.

--Mark Hill (talk) 19:15, 6 March 2016 (AEDT) added page Domenech-Mateu Collection.

--Mark Hill (talk) 07:11, 1 March 2016 (AEDT) updated Medicine lectures SH Lecture - Lymphatic Structure and Organs and SH Lecture - Respiratory System Development and practical Laboratory support Information.

Human Embryo (Carnegie stage 23) Palate



Older statistics have now been transferred to New 2016.


Older statistics have now been transferred to New 2015.


Older statistics have now been transferred to New 2014.

During 2014 the website had more than 18,000,000 total views, double that for the 2013 total views and more than the previous 3 years altogether.


Older statistics have now been transferred to New 2013

During 2013 the website had 9,016,452 total views compared to 11,763,978 in the 3 year period between 2009 to 2012.


Older statistics have now been transferred to New 2012.


Older statistics have now been transferred to New 2011.


Older statistics have now been transferred to New 2010.


Older statistics have now been transferred to New 2009.

Before 2009 Embryology was a website from 1996, rather than the current Wiki format.

Glossary Links

Glossary: A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z | Numbers | Symbols | Term Link

Cite this page: Hill, M.A. (2020, May 25) Embryology New. Retrieved from

What Links Here?
© Dr Mark Hill 2020, UNSW Embryology ISBN: 978 0 7334 2609 4 - UNSW CRICOS Provider Code No. 00098G