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== Development of the organs of Vision, Audition, Equilibrium and Olfaction ==
=== © Dr Mark Hill (2009) ===
==== [../People.htm Acknowledgements] ====
== Introduction ==

{| class="prettytable"
| [[Image:eyeear.gif]]
| This section of notes is a introduction to development of senses (sight, smell, hearing and balance) through the development of the specialized sense organs (eye, nose and ear). Portions of the eye, nose and ear appear very early in development as specialized surface regions ('''placodes''') on the embryo, these regions must be connected to the central nervous system by neural pathways that originate as extensions of the central nervous system.

There are now more specific notes on each sensory system [ear.htm Hearing and Balance], [eye.htm Vision], [nose.htm Smell] and [tongue.htm Taste] development.

|}

Related Pages: [week4_3.htm Week 4 Sensory Placodes] | [ear.htm Hearing] | [eye.htm Vision] | [nose.htm Smell] | [tongue.htm Taste]

== Reading ==
* '''Human Embryology '''(2nd ed.) Larson Ch12: p375-409
* '''The Developing Human: Clinically Oriented Embryology '''(6th ed.) Moore and Persaud Ch19: p491-511
* Essentials of Human Embryology Larson Ch12: p252-272
* Before We Are Born (5th ed.) Moore and Persaud Ch20: p460-479
* Human Embryology, Fitzgerald and Fitzgerald
* Search [http://www.ncbi.nlm.nih.gov/PubMed/medline.html PubMed- Medline]

== Objectives ==
* Understand the early development of the eye, defining optic vesicles, optic stalks, optic fissure, optic cups, lens placodes and lens vesicles.
* Understand the development of the retina, ciliary body, iris, lens, choroid, sclera and cornea.
* Understand the development of external, middle and internal ear, including otic placode, otocyst, endolymphatic duct and semicircular canals.

== Computer Activities ==
=== UNSW Embryology: ===
* [http://embryology.med.unsw.edu.au/Notes/senses4.htm#top Stage 13/14 Embryo]
** [http://embryology.med.unsw.edu.au/Notes/senses3.htm#Pig%20eye Eye] | [http://embryology.med.unsw.edu.au/Notes/senses3.htm#Pig%20ear Ear] | [http://embryology.med.unsw.edu.au/Notes/senses3.htm#Pig%20nose Nose]
* [http://embryology.med.unsw.edu.au/Notes/senses4.htm#top Stage 22 Embryo]
** [http://embryology.med.unsw.edu.au/Notes/senses4.htm#eye Eye] | [http://embryology.med.unsw.edu.au/Notes/senses4.htm#EAR Ear] | [http://embryology.med.unsw.edu.au/Notes/senses4.htm#nose Nose]
* [http://embryology.med.unsw.edu.au/Notes/senses5.htm#top Selected High Power Stage 22]
** [http://embryology.med.unsw.edu.au/Notes/senses5.htm#eye Eye ]| [http://embryology.med.unsw.edu.au/Notes/senses5.htm#ear Ear]
* [http://embryology.med.unsw.edu.au/Notes/senses7.htm Hearing and Balance development]
* [http://embryology.med.unsw.edu.au/Notes/senses6.htm#table Components of the Eye]
* [http://embryology.med.unsw.edu.au/Notes/senses5.htm#WWW%20Histology Histology] '''also look at [http://embryology.med.unsw.edu.au/Notes/senseslink.htm WWW Links]
* [http://embryology.med.unsw.edu.au//Series.htm Serial Sections]
** [http://embryology.med.unsw.edu.au//wwwhuman/hipower/HumHP.htm Human Selected]([../wwwhuman/hipower/HumA/A5L.htm optic nerve], [http://embryology.med.unsw.edu.au//wwwhuman/hipower/HumA/A6L.htm optic retina], [http://embryology.med.unsw.edu.au//wwwhuman/hipower/HumA/A7L.htm lens])'''
** [http://embryology.med.unsw.edu.au//wwwhuman/LowSet/HeadSet.htm#A6-B3 Stage 22 Head Images]
** [http://embryology.med.unsw.edu.au//wwwpig/lowset/HeadSet.htm#A6-B3 Stage 13/14 Head Images]
** [http://embryology.med.unsw.edu.au//wwwhuman/System/HumSysN.htm Human Systems]

'''Embryo Images Unit:'''

Eye Development and Ear Development

[http://www.med.unc.edu/embryo_images/ Embryo Images Online]

Links below require Internet connection.

* [http://www.med.unc.edu/embryo_images/unit-eye/eye_htms/eyetoc.htm Eye Development]
** [http://www.med.unc.edu/embryo_images/unit-eye/eye_htms/eye001.htm Eye Fields-Optic Vesicle (Weeks 3-4)]
** [http://www.med.unc.edu/embryo_images/unit-eye/eye_htms/eye009.htm Optic Cup, Lens Vesicle, Choroid Fissure, Hyaloid Artery (Weeks 5-6)]
** [http://www.med.unc.edu/embryo_images/unit-eye/eye_htms/eye016.htm Cornea, Anterior Chamber, Pupillary Membrane, Lens, Retina (Weeks 7-8)]
** [http://www.med.unc.edu/embryo_images/unit-eye/eye_htms/eye022.htm Iris, Cilliary Body (Weeks 9-15)]
** [http://www.med.unc.edu/embryo_images/unit-eye/eye_htms/eye025.htm Eyelids (Weeks 8-10)]
* [http://www.med.unc.edu/embryo_images/unit-ear/ear_htms/eartoc.htm Ear Development]
** [http://www.med.unc.edu/embryo_images/unit-ear/ear_htms/ear001.htm Inner Ear]
** [http://www.med.unc.edu/embryo_images/unit-ear/ear_htms/ear012.htm Middle Ear]
** [http://www.med.unc.edu/embryo_images/unit-ear/ear_htms/ear014.htm External Ear]

== References ==
For more recent review and research articles see specific notes on each Sensory System [ear.htm Hearing] | [eye.htm Vision] | [nose.htm Smell] | [tongue.htm Taste]

Selected Lists of References from PubMed March 1999 search results are available for Department of Anatomy computers without internet access. Computers with internet access can search from either [http://embryology.med.unsw.edu.au/Notes/senses2.htm#PubMed Page 2] or [http://www.ncbi.nlm.nih.gov/PubMed/medline.html PubMed Internet Access]

'''PubMed March 1999[../Refer/senses/eye_rev.htm Eye Development Reviews] | [../Refer/senses/ear_rev.htm Ear Development Reviews] | [../Refer/senses/nose_rev.htm Nose Development Reviews] | [../Refer/senses/select.htm Selected Research Articles and Reviews]

== Developmental Overview ==
'''Note these images are not to scale. Clicking each image will show full embryo.'''

[[Image:CSt16.gif]][[Image:CSt17.gif]]'''Carnegie Stage 15'''

'''Carnegie Stage 16'''

'''Carnegie Stage 17'''

[[Image:CSt19.gif]][[Image:CSt20.gif]]'''Carnegie Stage 18'''

'''Carnegie Stage 19'''

'''Carnegie Stage 20'''

[[Image:CSt22.gif]][[Image:CSt23.gif]]'''Carnegie Stage 21'''

'''Carnegie Stage 22'''

'''Carnegie Stage 23'''

[[Image:mov.gif]]''' See [../Movies/CSthead.mov Movie of these Head images (179 Kb)]

== Glossary ==
'''Use the alphabetical list below to find definitions of terms that are new to you or use the Google search window to search UNSW Embryology site.'''

{| class="prettytable"
| [Index/A.htm A] | [Index/B.htm B] | [Index/C.htm C] | [Index/D.htm D] | [Index/E.htm E] | [Index/F.htm F] | [Index/G.htm G] | [Index/H.htm H] | [Index/I.htm I] | [Index/J.htm J] | [Index/K.htm K] | [Index/L.htm L] | [Index/M.htm M] | [Index/N.htm N] | [Index/O.htm O] | [Index/P.htm P] | [Index/Q.htm Q] | [Index/R.htm R] | [Index/S.htm S] | [Index/T.htm T] | [Index/U.htm U] | [Index/V.htm V] | [Index/W.htm W] | [Index/X.htm X] | [Index/Y.htm Y] | [Index/Z.htm Z]
|

|}
== Quick Links ==

{| class="prettytable"
|
|
|
|

|}
=== UNSW Embryology ISBN: 978 0 7334 2609 4 ===
==== UNSW CRICOS Provider Code No. 00098G ====
[[Image:chris.gif]]

''' '''

=== Navigation ===
[../embryo.htm Home Page][../wwwhuman/Stages/Stages.htm Embryo stages][../wwwhuman/Fetus.htm Fetal Dev][../devnote.htm Dev Notes 1][../devnote2.htm Dev Notes 2][../sysnote.htm System Notes][../People.htm Acknowledgements]

=== Internal Links Senses ===
[http://embryology.med.unsw.edu.au/Notes/senses.htm Introduction][http://embryology.med.unsw.edu.au/Notes/senses2.htm#abnormal Abnormalities][http://embryology.med.unsw.edu.au/Notes/senses2.htm#OMIM OMIM][http://embryology.med.unsw.edu.au/Notes/senses2.htm#Questions Questions][http://embryology.med.unsw.edu.au/Notes/senses2.htm#PubMed Medline][http://embryology.med.unsw.edu.au/Notes/senses3.htm#Pig Pig Stage 13/14][http://embryology.med.unsw.edu.au/Notes/senses4.htm#Human Human (Stage22)][http://embryology.med.unsw.edu.au/Notes/senses5.htm Selected Human highpower][placode.htm Sensory Placodes][ear.htm Hearing][eye.htm Vision][tongue.htm Taste][http://embryology.med.unsw.edu.au/Notes/sensestxt.htm Text only page][http://embryology.med.unsw.edu.au/Notes/senseslink.htm WWW Links]

=== Internal Links ===
[../Movies/Movies.htm Movies][../Podcast/podcast.htm Audio][../News/news.htm News][../class.htm Class Notes][../Series.htm Serial Images][../K12/embryo.htm K12 Students][../Defect/page1.htm Abnormal][../histo.htm Histology][../sitemap.htm Site Map][Index/Index.htm Glossary][../Refer/refer.htm References][../Refer/search.htm Search]

=== External Links ===
[http://medicalsciences.med.unsw.edu.au/medsciences.nsf School of Med Sci][http://www.med.unsw.edu.au/ UNSW Medicine][http://www.unsw.edu.au/ University of NSW][http://cellbiology.med.unsw.edu.au/ UNSW Cell Biology][http://www.ncbi.nlm.nih.gov/books NCBI Books][http://www.ncbi.nlm.nih.gov:80/books/bv.fcgi?call=bv.View..ShowTOC&rid=dbio.TOC&depth=2 Dev Biol (Gilbert)][http://www.ncbi.nlm.nih.gov:80/books/bv.fcgi?call=bv.View..ShowTOC&rid=dbio.TOC&depth=2 2009-10-01T12:38:01Search Google]

<center>[[Image:Logo_PoweredBy_25wht.gif]]''' '''</center>

<center>''' '''</center>

<center><center>Search Google</center><center>Search NCBI</center>''' '''</center>

=== Search NCBIPage Details ===
==== [mailto:m.hill@unsw.edu.au Email]Copyright: Dr Mark HillCreated: 01.06.1997Updated:#BeginDate format:Ge1 17.07.2009#EndDate UNSW CRICOS Provider Code No. 00098G ====
=== Comments ===
[[Image:mhicon08.jpg]]

'''These notes introduce the development of the senses. Our senses are a combination of several different embryonic tissues, which for us to "sense" must also connect with the central nervous system.'''

'''This general section of sensory notes provides a brief introduction. More detailed information can be found on the specific sensory sysstem pages [ear.htm Hearing] | [eye.htm Vision] | [nose.htm Smell] | [tongue.htm Taste]. '''

<center>[[Image:retsm.gif]]''' '''</center>

<center>'''the retina by Santiago Ram√≥n y Cajal (c1900)'''</center>

Talk:2009 Lab 9 - Revision history

S8600021: Created page with '== Development of the organs of Vision, Audition, Equilibrium and Olfaction == === © Dr Mark Hill (2009) === ==== [../People.htm Acknowledgements] ==== == Introduction == {| cl…'